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Protein post-translational modification by ubiquitination participates in many aspects of plant growth,development and stress responses.Covalent attachment of ubiquitin to targets is sequentially catalyzed by three enzymes(E1,E2 and E3),among them the E2(Ubiquitin Conjugation enzyme,UBC)functions importantly in assisting E3(Ubiquitin ligase)for substrate specificity.