Hop(Humulus lupulus L.)is an important ingredient,along with barley and yeast,for the beer brewing industry.Hops glandular trichomes uniquely produce two important classes of secondary metabolites in abundance: bitter acids and xanthohumol,which give beer its flavor and have been used as potential anticancer drugs and nu-traceutical supplements.For these reasons,many studies had been performed on bitter acids and xanthohumol biosynthesis in the hop trichomes.