Plants host distinct bacterial communities on and inside various plant organs,of which those associated with the interactions between roots and microbes for nutrient availability,growth promotion,and disease suppression.The phylogenetic composition of these communities is defined by relatively few bacterial phyla,including Actinobacteria,Bacteroidetes,Firmicutes,and Proteobacteria.16S rDNA High-throughput sequencing approaches have provided recent insights into wetland plants,here we present a detailed characterization of the microbiomes of the Typha by deep sequencing,The spatial resolution of the study distinguished two compartments,the endosphere(organs and tissues interior),and rhizosphere(soil close to the root surface),each of which was found to harbor a distinct microbiome.A synthesis of available data suggests the bacterial microbiota of rhizosphere is differentiated from the endophytic bacteria.Diversity index(αandβ),Venn diagram as well as compositions of spatial compartments image support a multistep model for microbiome assembly from rhizosphere soil to endosphere wherein the root plays a selective gating role.Both endosphere and rhizosphere contain bacteria that provide indirect pathogen protection,but rhizobacteria members appear to serve additional host functions through the acquisition of nutrients from soil for plant growth.In this study,rhizosphere niches and environmental features were two factors contributing to microbiome variation.Similarities in the distribution of phyla in the microbiomes of Typha and other plants suggest that conclusions derived from this study might be generally applicable to wetland plants.