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Helicases are classified to SF1-SF6 super families, which are conserved in allspecies.DEAD-box RNA helicase belongs to the SF2 and is identified relative well in yeast and animal.In Arabidopsis there are 58 DEAD-box RNA helicase members, with functions mostly unknown.FAC1O encoded a putative DEAD-box RNA helicase which shared 56% identity with Has1p that is required for 40S ribosomal subunit biogenesis in Saccharomyces cerevisiae and 55% identity with Ddx18 that is essential for cell-cycle progression in human.