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Chaperonins are a class of molecular chaperones that assist in the folding and assembly of a wide range of substrates.In plants,chloroplast chaperonins are composed of two different types of subunits,Cpn60α and Cpn60β,and duplication of Cpn60α and Cpn60β genes occurs in a high proportion of plants.However,the importance of multiple Cpn60α and Cpn60β genes in plants is poorly understood.