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As one of the most important food sources for human,rice endosperm is composed of an outer aleurone and an inner starchy endosperm(white rice).It has been proposed that,in the caryopsis,sucrose delivered by phloem from photosynthetic leaves passes the maternal tissues through plasmadesmata,and then actively through aleurone to the developing endosperm.However,the exact role of aleurone in regulating sugar transport remains unknown.