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The vacuole in plant cell plays a crucial role in plants tolerance to salinity.Two plant proton pumps,vacuolar H+-ATPase (V-H+-ATPase) and H+-pyrophosphatase (V-H+-PPase),participate in acidifying compartments of the vacuoles,which establishes an electrochemical H+-gradient to drive sequestration of Na+ into the vacuole lumen,compartmentalizing this toxic ion from the cytoplasm and maintaining low cytoplasmic Na+ concentrations.