Homologs of LFY gene encode plant-specific transcription factors which play an important role in flower development.A wheat FLORICAULA/LEAFY ortholog,WFL,was reported to be associated with spikelet formation as lateral branch of the inflorescence meristem (Shitsukawa et al.,2006).Bread wheat is derived from a cross between emmer and Aegilops tauschii which took place some 8000 years ago.A better understanding of WFLs role from genetic and genomic perspectives will shed light on its natural evolution and its history under domestication.