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Sterols are membrane lipids crucial for cellular functions and development of diverse organisms.SQUALENE SYNTHASE (SQS) catalyzes the first committed step of sterol biosynthesis.In Arabidopsis there are two annotated SQS-coding genomic sequences, SQS1 and SQS2, organized in a tandem array.Previous reports showed that SQS2 has no SQS activity, SQS1 is the only functional SQS in Arabidopsis.