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芡实（ＥｕｒｙａｌｅｆｅｒｏｘＳａｌｉｓｂ．）花粉的败育主要发生在小孢子母细胞减数分裂时期和小孢子单核期与二核期。形成的不正常四分体中有一些具有不完整的横隔壁，一些则完全不形成横隔壁，而发育成具４个核的原生质团；气候异常是造成该时期不育现象的主要原因之一。小孢子单核期及二核期存在发育异常的孢子。绒毡层的提前发育、过早解体、肥大生长及延迟退化是花粉败育的一个重要原因。
The abortion of pollen of Euryale ferox Salisb. Mainly occurred during meiosis of microspore mother cell and microspore mononuclear and dinuclear phases. Some of the abnormal tetrad formed with incomplete transversal barrier, while others did not form transversal barrier at all and developed into 4 nuclear protoplasm. Abnormal climate was one of the main causes of infertility in this period . Microspores mononuclear and dinuclear abnormal development of spores. Advances in tapetal development, premature disintegration, hypertrophy and delayed degeneration are an important reason for pollen abortion.