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对原始椴树红松林中的灌木层及主要灌木种（毛榛子、溲疏和刺五加）的生物量、养分量、存留量、归还量、吸收量、循环速率等方面在林分的养分循环中的地位与作用进行了研究。研究结果表明：灌木层虽在林分的养分循环中处于次要地位，但具有促使林分养分步入良性生物循环的重要作用。毛榛子和溲疏是该林分灌木中的优势或亚优势种，在养分循环中起着重要的或突出的作用，是重点保护的灌木种。刺五加在养分循环中存留养分比较多，归还养分比较少，虽然消耗地力却是具有药用的灌木种，对它应既维持又要控制。
The biomass, nutrient content, storage, return, uptake, and circulation rate of the shrub layer and the main shrub species (hazelnuts, Shizhu and Acanthopanax senticosus) The status and role of the cycle were studied. The results show that although the shrub layer plays a secondary role in the nutrient cycling of the stands, it has the important role of promoting the stand nutrients into the benign biological cycle. Hair hazelnuts and shu sparse are the dominant or subdominant species in the shrubs. They play an important or prominent role in the nutrient cycle and are the shrub species under the protection. Acanthopanax most nutrient retention in the nutrient cycle, the return of nutrients is relatively small, although the consumption of fertility is medicinal shrub species, it should both maintain and control.