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Walsh 于1864年首先报道一种害虫有形态相似而致害性(virulence)不同的种群。生物型(biotype)一词,乃是1903年由 Johannsen 作为种(species)以下的一个分类单位而提出。现代昆虫学上所称的生物型是抗虫育种先驱 Painter 于三十年代定名的。害虫新生物型的出现是由抗虫品种诱发的,正像抗病品种导致病原菌产生新的生理小种(race)那样。植病学家 Flor 于1959年最早提出了一种设想:寄主植物每有一个主效的抗
Walsh first reported in 1864 that there were morphologically similar and virulence-diverse populations of pests. The word biotype was proposed by Johannsen in 1903 as a taxon below species. The so-called biotype in modern entomology is Painter’s pioneer in insect breeding, which was named in the thirties. Emergence of the pest neoplasm is induced by insect-resistant varieties, just as the disease-resistant cultivars cause the pathogen to produce new physiological races. Florist first proposed in 1959 a concept: the host plant for each one of the main effect of anti-