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重新研究Kobayashi(1968)报道的来自泰国的Nippononaia carinata,以及Delvene和Munt(2011)报道的来自西班牙的Subnippononaia fordi标本;建立新亚属Nippononaia(Thainippononaia)Stiller and Chen,以及新种Nippononaia(Thainippononaia)hispaniensis Stiller and Chen,并认为新亚属代表Nippononaia属内的最早演化阶段,填补了Nippononaiidae科演化史的一个空白,即Eonippononaia阶段与Nippononaia(Arctonaia)阶段之间的空白。
Re-study of Nippononaia carinata from Thailand reported by Kobayashi (1968) and Subnipponona fordii from Spain reported by Delvene and Munt (2011); establishment of a new subgenus Stiller and Chen from Nippononaia (Thainippononaia) and a new species of Nippononaia (Thainippononaia) hispaniensis Stiller and Chen argues that the earliest evolutionary stage within the genus Nippononaia of the new subgenus fills a gap in the evolutionary history of the Nippononaiidae family, the gap between the Eonippononaia and Nippononaia (Arctonaia) stages.