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对平流层辐射处理ＳＰ３谷子和对照ＣＫ３谷子雄性细胞发育的研究表明，雄性细胞正常发育过程从孢原细胞开始，经初生造孢细胞、次生造孢细胞、小孢子母细胞、二分体、四分体和单核小孢子中央期、单核小孢子液泡期（单核靠边期）、单核小孢子成熟期直到二细胞花粉、三细胞成熟花粉结束。其中四分体形成属连续型。雄性细胞异常发育有几种情况：小孢子母细胞强烈液泡化，细胞质收缩解体，不能进入减数分裂；小孢子母细胞液泡化，虽能进入减数分裂，但不能形成正常二分体而中途退化；二分体、四分体细胞液泡化，不能进行正常发育；单核小孢子中央期，细胞质收缩包围核，小孢子形状扭曲呈月牙形或不规则形，最终细胞质和核解体而呈月牙形的空壳小孢子；单核液泡期的小孢子不能积聚营养物质，细胞质解体而成为大的空壳花粉粒。平流层辐射ＳＰ３谷子雄性细胞异常发育的比率达５０％左右，这可能与平流层辐射引起的染色体畸变以及ＳＰ１、ＳＰ２谷子染色体分离、重组有关。
Studies on the development of male stratum cells in the stratosphere of SP3 and control CK3 show that the normal development of male cells begins with the sporogenous cells and the primary cells, secondary sporogenous cells, microspore mother cells, The tetrad and monocyte microspore central stage, the mononuclear microspore vacuolar stage (mononuclear marginal stage), the mononuclear microspore maturation stage until the two-cell pollen and the three-cell mature pollen end. Which form a continuous tetrad. There are several conditions for the abnormal development of male cells: the microspore mother cells strongly vacuolize and the cytoplasm shrinks and disintegrates and can not enter meiosis. Although the microspore mother cells are vacuolized, they can enter meiosis but can not form normal doubles and go halfway Degenerate; bisected, tetrad cell vacuoles, can not be normal development; mononuclear microspore central phase, the cytoplasm shrinks around the nucleus, the shape of microspore twisted crescent or irregular shape, the final cytoplasm and nuclear disintegration was crescent Shaped hollow microspores; microspore of mononuclear vacuolar stage can not accumulate nutrients, cytoplasm disintegration and become a large shell pollen grains. Stratospheric radiation SP3 millet cells abnormal development rate of about 50%, which may be related to chromosome aberrations caused by stratospheric radiation and SP1, SP2 millet chromosome segregation, restructuring.