组3. 翠雀花组 Sect. Delphinastrum

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  Sect. Delphinastrum DC. Syst. 1: 351. 1818; et Prodr. 1: 53. 1824; W. T. Wang in Acta Bot. Sin. 10(1): 155. 1962; Tamura in Sei. Rep. Osaka Univ. 15: 33. 1966; et in Hiepko, Nat. Pflanzenfam., Zwei. Aufl., 17aIV: 299. 1995; W. T. Wang & M. J. Warnock in Fl. China 6: 246. 2001. Lectotype: Delphinium grandiflorum L. (Wang, 1962).
  Sect. Schizopetala Huth in Bull. Herb. Boiss. 1: 333. 1893.
  Sect.Diedropetala Huth in Bot. Jahrb. 20: 420. 1895. — Gen. Diedropetala (Huth) Galushko in Novost. Syst. Pl. Vasc. 15: 251. 1976.
  Sect. Kalobopetala Huth in l. c. 458
  Sect. Homochroe Bruhl in Ann. Bot. Gard. Calc. 5: 91. 1896.
  Sect. Isocheilae Bruhl in l. c., p. p.
  不具块根。叶3浅裂至3全裂。花序总状或伞房状,或花单朵顶生。萼片脱落,蓝色或紫色,稀白色,多少展开,稀上萼片船形;距与萼片近等長或比萼片长,稀比萼片短。花瓣与萼片同色,稀淡黄色或黄白色。退化雄蕊瓣片,也与萼片同色,稀淡黄色或黄白色,2浅裂或2半裂,稀不分裂,腹面基部之上被黄色或白色髯毛。心皮3,稀5。种子四面体形,沿纵棱具狭翅或无翅,或狭倒卵球形,具密集横向排列的小鳞片。
  约190种,隶属于6亚组,广布于亚洲、欧洲东北部、非洲和北美洲。我国有6亚组约132种,多数种分布于西南部的四川、云南、西藏、青海等省(区),西部、北部和东北部诸省各有4种至1种。
  分种和变种检索表
  1. 花瓣和退化雄蕊淡黄色或淡黄白色;萼片蓝色或紫色。
  2. 叶片3全裂;花单朵顶生;萼片比距长;花瓣淡黄白色;退化雄蕊瓣片淡黄白色,不分裂;心皮5
  159. 湟中翠雀花Delphinium huangzhongense
  2. 叶片3深裂;花组成总状花序;退化雄蕊瓣片2浅裂;心皮3或1。
   3. 萼片与距近等长;花瓣和退化雄蕊淡黄色;心皮3。
  4. 花较小,组成复总状花序;萼片长6~7 mm,顶端以直角向内弯曲;心皮无毛
  160. 征镒翠雀花 D. zhengyianum
  4. 花较大,组成简单总状花序;萼片长14 mm,不弯曲;子房被短柔毛
  161. 文采新翠雀花 D. neowentsaii
   3. 萼片比距短;花瓣和退化雄蕊黄白色;心皮1,无毛
  189. 单蕊翠雀花 D. monogynum
  1. 花瓣和退化雄蕊与萼片同色,即呈蓝色或紫色,稀白色。
  5. 基生叶多不存在,或存在,则在花期枯萎。
   6. 种子具3纵棱,沿棱具狭翅。
  7. 心皮4~6
   8. 叶片3全裂;心皮5
  98. 密叶翠雀花 D. kingianum
  9. 茎有毛;叶全裂片顶端急尖。
   10. 茎密被白色短柔毛和黄色短腺毛。
  11. 子房密被短柔毛
  98a. 模式变种 var. kingianum
  11. 子房只在腹缝线上有毛,其他部分无毛
  98b. 光果密叶翠雀花 var. leiocarpum
   10. 茎密被白色短柔毛,具少数黄色腺毛
  98c. 少腺密叶翠雀花 var. eglandulosum
  9. 茎无毛;叶全裂片顶端长渐尖
  98d. 尖裂密叶翠雀花 var. acuminatissimum
   8. 叶片3深裂;心皮4~6
  99. 乡城翠雀花 D. humilius
  7. 心皮3。
   12. 叶片3全裂;萼片比距长
  100. 尾裂翠雀花 D. caudatolobum
   12. 叶片3深裂;萼片与距近等长,或比距短。
  13. 总状花序轴和花梗被白色短柔毛和黄色短腺毛
  103. 拉萨翠雀花 D. gyalanum
  13. 总状花序轴和花梗无黄色腺毛。
   14. 总状花序轴和茎无毛。
  15. 植株高达140 cm;萼片长14~15 mm;退化雄蕊瓣片2浅裂
  106. 唇花翠雀花 D. cheilanthum
   16. 花梗被白色反曲短柔毛
  106a. 模式变种 var. cheilanthum
   16. 花梗被黄色伸展短毛
  106b. 展毛唇花翠雀花 var. pubescens
  15. 植株高达20~30 cm;萼片长15~20 mm。
   17. 叶侧生1回裂片2半裂;萼片长15~20 mm,距长18~24 mm;退化雄蕊瓣片近圆形, 2浅裂   104. 宽片翠雀花 D. latilimbum
   17. 叶侧生1回裂片2深裂超过叶片中部;萼片长15~18 mm,距长20 mm;退化雄蕊瓣片宽椭圆形,顶端微凹
  107. 毓泉翠雀花 D. yuchuanii
   14. 花梗和茎有毛,稀茎近无毛 (D. hsinganense)。
  18. 萼片两面均有毛,与距等长或稍长
  101. 粗距翠雀花 D. pachycentrum
   19. 萼片长圆形
  101a. 模式变种var. pachycentrum
   19. 萼片披针形
  101b. 狭萼粗距翠雀花var. lancisepalum
  18. 萼片内面无毛。
   20. 花序复总状;小苞片贴伏于花上,狭披针形,长6~11 mm;萼片长13 mm;退化雄蕊瓣片倒卵状长圆形,有白色髯毛
  102. 拟粗距翠雀花D. pachycentroides
   20. 花序简单总状;小苞片与花离开,钻形,长3~5 mm;萼片长14~17 mm;退化雄蕊瓣片圆卵形,有黄色髯毛
  105. 兴安翠雀花 D. hsinganense
   6. 种子具密集横向排列的小鳞片或横波状条纹。
  21. 叶片宽心形,不分裂或不明显3浅裂
  158. 心叶翠雀花 D. calthifolium
  21. 叶片五角形或肾形,3浅裂,3深裂或3全裂。
   22. 叶片3浅裂或3深裂。
  23. 叶片3浅裂,分裂不超过叶片中部。
   24. 叶片3浅裂不达叶片中部,浅裂片互相不覆压;总状花序长10 cm;花梗被白色柔毛和黄色腺毛
  135. 葡萄叶翠雀花 D. sinovitifolium
   24. 叶片3浅裂达叶片中部,浅裂片互相覆压;伞房花序长6 cm;花梗无毛
  134. 茂县翠雀花 D. maoxianense
  23. 叶片3深裂超过中部,达近基部。
   25. 花序伞房状。
  26. 茎无毛。
   27. 萼片背面顶端之下具角状突起
  133. 叉角翠雀花 D. furcatocornutum
   27. 萼片无角状突起。
  28. 萼距螺旋状弯曲
  137. 川黔翠雀花 D. bonvalotii
  28. 萼距稍向下弯曲
  130. 毛梗翠雀花 D. eriostylum
  26. 茎有毛。
   29. 茎被短柔毛;叶片宽达6 cm
  131. 平武翠雀花 D. pingwuense
   29. 茎被糙硬毛;叶片宽达12 cm。
  30. 花梗有黄色腺毛
  130. 毛梗翠雀花D. eriostylum
   31. 花梗只被黄色腺毛;子房疏被柔毛
  130a. 模式变种var. eriostylum
   31. 花梗被黄色短腺毛和白色短柔毛;心皮无毛
  130b. 糙叶毛梗翠雀花 var. hispidum
  30. 花梗无黄色腺毛
  132. 疏伞翠雀花D. laxicymosum
   32. 花梗无毛;萼距长23~27 mm
  132a. 模式变种var. laxicymosum
   32. 花梗被短柔毛;萼距长20 mm
  132b. 毛序疏伞翠雀花var. pilostachyum
   25. 花序总状。
  33. 萼距直,稍向下弯曲,或成直角向下弯曲。
   34. 萼片比距长;花梗长5~9 cm
  108. 长梗翠雀花D. longipedicellatum
   34. 萼片与距近等长或较短。
  35. 萼片与距近等长。
   36. 萼距向斜上方伸展,长10~13 mm
  110. 直距翠雀花D. orthocentrum
   36. 萼距不向上方伸展。
  37. 茎被短柔毛;萼距水平方向伸展,长12~16 mm。
   38. 花梗有少数黄色腺毛;萼片长12~18 mm,距长10~16 mm;子房被短柔毛
  111. 小瓣翠雀花D. micropetalum
   38. 花梗无黄色腺毛;萼片长12 mm,距长9~12 mm;心皮无毛
  109. 拟直距翠雀花D. orthocentroides
  37. 茎被长硬毛;萼距稍向下弯曲,长20 mm
  112. 毛茎翠雀花D. hirticaule
   39. 花梗无毛
  112a. 模式变种var. hirticaule
   39. 花梗有黄色腺毛
  112b. 腺毛毛茎翠雀花var. mollipes
  35. 萼片比距短。
   40. 小苞片与花邻接或近邻接,并贴伏于花上。
  41. 小苞片狭条形
  122. 陰地翠雀花D. umbrosum
   42. 小苞片宽0.5~1.1 mm。
  43. 茎高达110 cm,被短糙伏毛;小苞片宽0.5~0.6 mm
  122a. 模式变种var. umbrosum   43. 茎高达70 cm,被开展硬毛;小苞片宽0.6~1.1 mm
  122b. 展毛阴地翠雀花var. hispidum
   42. 小苞片宽1.5~1.8 mm
  122c. 宽苞阴地翠雀花 var. drepanocentrum
  41. 小苞片条状披针形。
   44. 茎被短糙伏毛;萼片外面被短柔毛
  120. 滇川翠雀花 D. delavayi
  45. 花梗被白色短糙伏毛和黄色短腺毛
  120a. 模式变种var. delavayi
  45. 花梗只被白色短糙伏毛
  120b. 保山翠雀花var. baoshanense
   44. 茎密被开展糙硬毛;萼片外面疏被糙硬毛
  121. 须花翠雀花D. pogonanthum
  46. 雄蕊无毛
  121a. 模式变种var. pogonanthum
  46. 雄蕊花丝每侧有1~2根毛
  121b. 毛蕊须花翠雀花var. lasiandrum
   40. 小苞片多少与花分离,不贴伏花上。
  47. 叶中央裂片不明显3浅裂或不分裂。
   48. 叶中央裂片菱形,边缘中部之上具牙齿,中部之下全缘。
  49. 萼片长9 mm,距长17 mm;茎被糙伏毛或无毛
  113. 淡紫翠雀花D. handelianum
  49. 萼片长15~16 mm,距长20~21 mm;茎无毛
  114. 河南翠雀花 D. honanense
   50. 花梗被白色短柔毛和黄色短腺毛
  114a. 模式变种var. honanense
   50. 花梗无黄色腺毛
  114b. 毛梗河南翠雀花 var. piliferum
   48. 叶中央裂片披针形,自基部之上至顶端有密集牙齿
  115. 毛叶翠雀花D. hirtifolium
  47. 叶中央裂片菱形,近中部明显3浅裂,稀3深裂。
   51. 叶中央裂片宽菱形,3深裂近中脉
  129. 假深蓝翠雀花D. pseudocyananthum
   51. 叶中央裂片菱形,3浅裂。
  52. 茎无毛。
   53. 花梗无毛
  136. 黑水翠雀花D. potaninii
  54. 心皮无毛;蓇荚直,长圆形,长1.4~1.7 cm。
   55. 小苞片宽0.2~0.3 mm
  136a. 模式变种var. potaninii
   55. 小苞片宽1~1.2 mm
  136b. 宽苞黑水翠雀花var. latibracteolatum
  54. 心皮有疏毛;蓇荚新月形,长0.5~1 cm
  136c. 九峰山翠雀花var. jiufengshanense
   53. 花梗有毛。
  56. 花梗密被黄色短腺毛
   57. 总状花序复杂;心皮无毛。
  58. 草本高1.5~2 m;花梗被白色短糙伏毛和黄色短腺毛;萼片长11 mm,距长18 mm;退化雄蕊瓣片2裂超过其中部
  125. 多花翠雀花D. polyanthum
  58. 草本高約80 cm;花梗只被黄色短腺毛;萼片长7~10 mm,距长11~12 mm;退化雄蕊瓣片顶端微凹
  126. 左贡翠雀花D. zuogongense
   57. 总状花序简单;子房被柔毛
  127. 川西翠雀花D. tongolense
  59. 叶柄无毛
  127a. 模式变种var. tongolense
  59. 叶柄被长硬毛
  127b. 毛柄川西翠雀花var. hirsutipes
  56. 花梗只被白色短柔毛,无黄色腺毛;总状花序复杂;草本高约1 m
  128. 高茎翠雀花D. altissimum
  52. 茎有毛。
   60. 茎和花梗被白色短柔毛和黄色短腺毛
  123. 米林翠雀花 D. sherriffii
   60. 茎和花梗无黄色腺毛。
  61. 茎被开展糙硬毛;退化雄蕊瓣片被黄色髯毛。
   62. 总状花序简单;萼片外面被白色短柔毛和黄色短腺毛;退化雄蕊的爪细,比瓣狭约5倍
  116. 峨眉翠雀花 D. omeiense
  63. 萼片长12~16mm,距长20~26 mm
  116a. 模式变种 var. omeiense
  63. 萼片长4~8mm,距长7~18mm
  116b. 小花峨眉翠雀花 var. micranthum
   62. 总状花序复杂;萼片外面只疏被黄色腺毛,无白色毛,长10 mm,距长20 mm;退化雄蕊的爪条形,比瓣片稍狭
  119. 宽爪翠雀花 D. platyonychium
  61. 茎被柔毛。
   64. 茎被开展柔毛;总状花序复杂;萼距长23 mm
  124. 盐源翠雀花 D. yanyuanense
   64. 茎被反曲短柔毛;总状花序简单;萼距长17~18 mm。
  65. 叶片3深裂至距基部2~4 mm处; 上萼片狭卵形, 13 × 6 mm; 退化雄蕊瓣片被黄色髯毛   117. 柔毛翠雀花 D. pubescens
  65. 叶片3深裂至距基部6~7 mm处; 上萼片宽卵形,10 × 7 mm; 退化雄蕊瓣片被白色髯毛
  118. 周城翠雀花 D. zhouchengense
  33. 萼距向下成U字形或螺旋状弯曲。
  66. 茎无毛。
   67. 花梗无毛
  136. 黑水翠雀花 D. potaninii
   67. 花梗有毛。
  68. 花梗只被黄色短腺毛;叶片宽达16 cm,3深裂稍超过中部;小苞片条形
  141. 汶川翠雀花 D. wenchuanense
  68. 花梗被白色短柔毛和黄色短腺毛;叶片宽达12 cm,3深裂至近基部;小苞片丝形。
   69. 小苞片长4.5~7 mm;萼距长12~21 mm
  140. 拟川西翠雀花 D. pseudotongolense
   69. 小苞片长1.5~4 mm;萼距长18~24 mm
  138. 拟螺距翠花 D. bulleyanum
  70. 子房疏被柔毛
  138a. 模式变种 var. bulleyanum
  70. 心皮无毛
  138b. 光果拟螺距翠雀花 var. leiogynum
  66. 茎有毛。
   71. 心皮无毛
  139. 贡嘎翠雀花 D. gonggaense
   71. 子房有毛。
  72. 茎被开展硬毛;叶片宽达10(~14) cm;萼片长15~18 mm
  142. 螺距翠雀花 D. spirocentrum
  72. 茎被开展短柔毛;叶片宽达5.4 cm;萼片长12~13 mm
  143. 狭菱形翠雀花 D. angustirhombicum
  22. 叶片3全裂。
  73. 叶全裂片细裂,末回裂片条状披针形
  157. 塔城翠雀花 D. aemulans
  73. 叶全裂片不细裂,末回裂片卵形或三角形。
   74. 萼距长15 mm以下。
  75. 茎、花梗和心皮均无毛
  144. 疏花翠雀花 D. sparsiflorum
  75. 茎和花梗有毛。
   76. 草本高约100 cm;总状花序复杂;萼距长8~9 mm
  145. 狭序翠雀花 D. angustipaniculatum
   76. 草本高达60~80 cm;总状花序简单;萼距长12~15 mm。
  77. 叶中央裂片菱形,在中部3浅裂,或近羽状分裂;子房被柔毛
  146. 错那翠雀花 D. conaense
  77. 叶中央裂片宽菱形,在中部之下3深裂;心皮无毛
  147. 秀丽翠雀花 D. wangii
   74. 萼距长20~28 mm。
  78. 茎无毛。
   79. 萼片无角状突起。
  80. 花梗无毛。
   81. 草本高达110 (~150) cm;叶片宽达20 cm;总状花序复杂;小苞片长2.5~3.5 mm;萼距直或向下弯曲
  151. 秦岭翠雀花 D. giraldii
   81. 草本高达70 cm;叶片宽达8 cm;总状花序简单;小苞片长4~5 mm;萼距螺旋状弯曲
  152. 光茎翠雀花 D. glabricaule
  80. 花梗有毛。
   82. 花梗被白色短柔毛和黄色短腺毛;总状花序简单
  148. 木里翠雀花 D. muliense
  83. 小苞片披针形或披针状条形,8~13 × 1.2~2 mm
  148a. 模式变种 var. muliense
  83. 小苞片近丝形,3.5~4 × 0.2 mm
  148b. 小苞木里翠雀花 var. minutibracteolatum
   82. 花梗只被白色短柔毛,无黄色腺毛;总状花序复杂
  153. 松潘翠雀花 D. sutchuenense
   79. 萼片外回顶端之下具角状突起。
  84. 花梗被白色短柔毛; 萼片角状突起极小, 不明显, 距水平伸展或稍向下弯曲; 心皮3
  153. 松潘翠雀花 D. sutchuenense
  84. 花梗被黄色短腺毛;萼片角状突起明显,长2.5~3.5,距呈U字形或螺旋状弯曲;心皮3(~4)
  154. 弯距翠雀花 D. campylocentrum
  78. 茎有毛。
   85. 花梗无毛。
  86. 叶片下面无毛;萼片无毛,距长28 mm
  156. 大藏翠雀花 D. dazangense
  86. 葉片下面有毛;萼片有毛,距长20 mm。
   87. 茎下部被黄色硬毛;萼片内面无毛
  149. 壤塘翠雀花 D. rangtangense
   87. 茎下部被白色硬毛;萼片内面有疏毛
  150b. 光序拟弯距翠雀花 D. pseudocampylocentrum var. glabripes
   85. 花梗有毛。
  88. 草本高50 cm;花梗疏被黄色短腺毛;小苞片与花分开;萼距长17~22 mm;子房有疏毛   150a. 拟弯距翠雀花 D. pseudocampylocentrum var. pseudocampylocentrum
  88. 草本高50~90 (~100) cm;花梗密被白色短柔毛;小苞片与花邻接;萼距长22~25 mm;心皮无毛
  155. 三小叶翠雀花 D. trifoliolatum
  5. 基生叶发育,稀不存在,或存在时在花期枯萎。
  89. 叶片3浅裂或3深裂。
   90. 种子近四面体形,沿纵棱具翅。
  91. 萼片外面顶端之下具角状突起。
   92. 叶片3裂稍超过中部,末回裂片卵形或宽三角,稀狭披针或披针状条形。
  93. 叶一回中央裂片3次分裂
  166. 大理翠雀花 D. taliense
   94. 叶末回裂片卵形或三角形。
  95. 萼距基部粗2~3 mm。
   96. 萼片外面被短柔毛。
  97. 茎无毛。
   98. 心皮无毛
  166a. 模式变种var. taliense
   98. 子房有毛
  166b. 毛蕊大理翠雀花 var. glabrum
  97. 茎下部密被短柔毛
  166c. 毛茎大理翠雀花 var. pubicaule
   96. 萼片外面被长硬毛
  166d. 硬毛大理翠雀花 var. hirsutum
  95. 萼距基部粗4~5 mm
  166e. 粗距大理翠雀花 var. platycentrum
   94. 叶末回裂片狭披针或披针状条形;萼距比萼片长2倍
  166f. 长距大理翠雀花 var. dolichocentrum
  93. 叶的1回中央裂片1~2次分裂
  167. 角萼翠雀花 D. ceratophorum
   99. 萼片角状突起长1~2 mm。
  100. 叶片宽达10 cm,末回裂片卵形或宽三角形。
   101. 萼片外面疏被短糙伏毛(毛长约0.4 mm),角状突起长1.5~2 mm
  167a. 模式变种 var. ceratophoram
   101. 萼片外面被长硬毛(毛长约2 mm),角状突起长1~1.5 mm
  167c. 毛角萼翠雀花 var. hirsutum
  100. 叶片宽8~19 cm,末回裂片条状披针形;萼片角状突起长1.5~2 mm
  167d. 粗壮角萼翠雀花 var. robustum
   99. 萼片角状突起长在1 mm以下
  167b. 短角萼翠雀花 var. brevicorniculatum
   92. 叶片3深裂至近叶片基部处。
  102. 花梗上部被黄色短腺毛和白色硬毛;萼片长10~12 mm;退化雄蕊瓣片被白色髯毛
  169. 短萼翠雀花 D. brevisepalum
  102. 花梗无毛;萼片长14~22 mm;退化雄蕊瓣片被黄色髯毛。
   103. 茎无毛;萼片长18~22 mm,角状突起长1 mm,距长24 mm
  168. 拟角萼翠雀花 D. ceratophoroides
   103. 茎下部被糙伏毛; 萼片长14~16 mm, 角状突起长1~1.5 mm, 距长27~28 mm
  170. 拟长距翠雀花 D. dolichocentroides
  91. 萼片无角状突起。
   104. 叶片3裂达或稍超过中部。
  105. 花梗无毛
  162. 钩距翠雀花 D. hamatum
  105. 花梗被白色短柔毛和黄色短腺毛。
   106. 茎近顶部被黄色短腺毛;萼片长9 mm,距长14~17 mm;心皮无毛
  163. 宽菱形翠雀花 D. latirhombicum
   106. 茎无毛;萼片长15~19 mm,距长20~22 mm;子房被柔毛
  165. 宁朗山翠雀花 D. ninglangshanicum
   104. 叶片3深裂至近基部处。
  107. 萼片长13~19 mm。
   108. 花梗无黄色腺毛。
  109. 花梗上部具2小苞片。
   110. 总状花序长2~6.5(~16)cm;子房密被柔毛。
  111. 总状花序长16 cm;花梗长3.5~10 cm
  174. 岩瓦翠雀花 D. yanwaense
  111. 总状花序长2~6.5 cm;花梗长1.4~3.2 cm。
   112. 萼距长25~27 mm;退化雄蕊瓣片2半裂,被白色髯毛
  172. 维西翠雀花 D. weixiense
   112. 萼距長18 mm;退化雄蕊瓣片顶端微凹,被黄色髯毛
  186. 洱源翠雀花 D. lankongense
   110. 总状花序长10~26 mm;子房只腹缝线有毛,其他部分无毛
  187. 密距翠雀花 D. pycnocentrum
  109. 花梗在顶端具2条形小苞片
  164. 拟阴地翠雀花 D. umbrosoides
   108. 花梗被黄色短腺毛和白色短柔毛;萼距长16~18 mm。   113. 茎无毛;雄蕊花丝有毛。
   113′. 总状花序长22 cm;退化雄蕊瓣片倒心形,宽7 mm;花药无毛
  182. 倒心片翠雀花 D. obcordatilimbum
   113′. 总状花序长12~15 cm;退化雄蕊瓣片长圆形,宽3 mm;花藥背面有毛
  179. 毛药翠雀花D. lasiantherum
  113. 茎有毛; 花丝无毛。
   113″. 茎被短柔毛,有1叶,基生叶约7;退化雄蕊瓣片2裂稍超过中部,被白色髯毛;子房被短柔毛
  171. 缺刻翠雀花D. incisolobulatum
   113″. 茎被糙伏毛,有数叶,基生叶在花期枯萎;退化雄蕊瓣片不分裂或2微裂,被黄色髯毛;心皮无毛
  173. 甘孜翠雀花D. kantzeense
  107. 萼片长20~30 mm。
   114. 叶柄鞘发育良好,宽8~15 mm
  180. 鞘柄翠雀花 D. coleopodum
   114. 叶柄鞘很狭窄或不存在。
  115. 花梗无毛。
   116. 茎被短柔毛;萼片长9~14 mm,距长17~24 mm;退化雄蕊瓣片2半裂
  178. 云南翠雀花 D. yunnanense
   116. 茎无毛;萼片长18 mm,距长26~30 mm;退化雄蕊瓣片不分裂
  185. 凉山翠雀花 D. liangshanense
  115. 花梗有毛。
   117. 花梗有黄色腺毛。
  118. 茎被硬毛;叶片两面疏被糙伏毛;萼距长17~20 mm
  175. 永宁翠雀花 D. yongningense
  118. 茎被短柔毛;叶片无毛;萼距长22 mm
  176. 纸叶翠雀花 D. pergameneum
   117. 花梗无黄色腺毛。
  119. 草本高3~8 cm;花梗被白色短柔毛;萼片长15~16 mm,距长25 mm
  181. 先骕翠雀花 D. hui
   120. 叶片的中裂片具3齿;茎顶端具1~2花
  181a. 模式变种var. hui
   120. 叶片的中裂片具3~5齿;茎顶端具1~5(~10)花
  181b. 多裂叶翠雀花var. lobulibundum
  119. 草本高11~18 cm;花梗被黄色短柔毛;萼距长28~29 mm。
   121. 茎下部疏被开展短柔毛;小苞片长5 mm;退化雄蕊瓣片顶端圆形,不分裂,全缘
  183. 宁蒗翠雀花 D. pseudohamatum
   121. 茎被反曲短柔毛;小苞片长(5~)7.5~9 mm;退化雄蕊瓣片顶端微凹或2微裂
  184. 玉龙山翠雀花 D. yulongshanicum
  90. 种子具密集横向排列的小鳞片; 叶片3裂稍超过中部; 萼片长13~15 mm, 距长20~22 mm
  188. 会泽翠雀花 D. hueizeense
  89. 叶片3全裂。
  122. 种子近四面体形,沿纵棱具狭翅。
   123. 花序总状。
  124. 上萼片船形。
   125. 叶片基部深心形,末回裂片三角形或狭披针形;萼距比萼片短,筒状钻形,长11~18 mm
  190. 丽江翠雀花 D. likiangense
   125. 叶片基部近截形,末回裂片条形;萼距比萼片长,近筒形,长21~27(~30)mm
  191. 川甘翠雀花 D. souliei
  124. 上萼片多少展开,不呈船形。
   126. 叶的末回裂片卵形,三角形或狭披针形。
  127. 萼片比距长或与距近等长。
   128. 茎和花梗被短柔毛;萼片比距长,长17~23 mm,距长12~16 mm
  192. 墨脱翠雀花 D. medogense
   128. 茎和花梗无毛;萼片和距均长10~12 m
  193. 德浚翠雀花 D. yuanum
  127. 萼片比距短。
   129. 茎密被开展短柔毛
  210. 威海翠雀花 D. weihaiense
   129. 茎被贴伏短柔毛或硬毛,或无毛。
  130. 花梗被黄色短腺毛和白色短柔毛
  204b. 锐裂翠雀花 D. thibeticum var. laceratilobum
  130. 花梗无黄色腺毛。
   131. 花梗无毛;叶均基生
  199. 短葶翠雀花 D. breviscaposum
   131. 花梗有毛。
  132. 茎和心皮无毛;萼距长15 mm
  194. 丝苞翠雀花 D. filibracteolatum
  132. 茎和心皮有毛。
   133. 退化雄蕊瓣片不分裂。
  134. 萼距长24~28 mm;退化雄蕊瓣片顶端啮蚀状
  201. 囊谦翠雀花 D. nangchienense
  134. 萼距长16~24 mm;退化雄蕊瓣片顶端微凹。
   135. 小苞片狭长圆形,长11~18 mm;萼距长达19 mm   195. 威宁翠雀花 D. weiningense
   135. 小苞片狭条形或钻形,长2.5~6 mm;萼距长达24 mm
  200. 金沙翠雀花 D. majus
   133. 退化雄蕊瓣片2裂。
  136. 叶均基生, 中央全裂片近楔形, 1回分裂, 近頂端2浅裂或具3牙齿
  198. 察隅翠雀花 D. chayuense
  136. 叶基生并茎生,中央全裂片2回分裂。
   137. 茎被反曲短柔毛;叶中央全裂片楔状菱形;小苞片长圆形,长9~12 mm;萼片长15~18 mm,距长14~20 mm
  196. 太白翠雀花 D. taipaicum
   137. 茎被开展硬毛; 叶中央全裂片菱形; 小苞片条形, 长7~8 mm; 萼片长10~14 mm, 距长20~24 mm
  197. 拟澜沧翠雀花 D. pseudothibeticum
   126. 叶的全裂片细裂,末回裂片披针状条形、条形或狭条形。
  138. 心皮5。
   139. 花序轴和花梗被反曲短柔毛, 无黄色腺毛; 萼片长9 mm, 距长12 mm; 退化雄蕊瓣片顶端微凹; 心皮无毛
  202. 五果翠雀花 D. sinopentagynum
   139. 花序轴和花梗有黄色腺毛; 萼片长15 mm, 距长20~24 mm; 退化雄蕊瓣片2微裂; 子房被柔毛
  203. 条裂翠雀花 D. pseudomosoynense
  140. 花序轴和花梗被白色短柔毛和黄色短腺毛;上萼片外面顶端之下有1小角状突起,距长20 mm;花瓣顶端被缘毛
  203a. 模式变种 var. pseudomosoynense
  140. 花序轴和花梗只被黄色短腺毛, 无白色短柔毛; 上萼片无角状突起, 距长21~24 mm; 花瓣无毛
  203b. 疏毛条裂翠雀花 var. subglabrum
  138. 心皮3。
   141. 退化雄蕊瓣片长圆形成倒卵状长圆形。
  142. 花序轴和花梗无毛。
   143. 叶末回裂片宽6 mm; 总状花序长25~45 cm; 萼距长13~15 mm; 退化雄蕊瓣片长4.6 mm, 2半裂
  177. 拟云南翠雀花 D. pseudoyunnanense
   143. 叶末回裂片宽1.6~3 mm;总状花序长10~20 cm;萼距长25~28 mm;退化雄蕊长8 mm,2微裂
  205. 光轴翠雀花 D. leiostachyum
  142. 花序轴和花梗有毛。
   144. 花序轴和花梗只被白色短柔毛;小苞片与花分开
  208. 孟获营翠雀花 D. mosoynense
   144. 花序轴和花梗被白色短柔毛和黄色短腺毛;小苞片与花邻接
  204a. 澜沧翠雀花 D. thibeticum var. thibeticum
   141. 退化雄蕊瓣片宽倒卵形或近圆形。
  145. 茎和花梗被短柔毛和短腺毛。
   146. 茎和花梗密被开展白色短柔毛和白色短腺毛
  211. 烟台翠雀花 D. chefoense
   146. 茎和花梗密被贴伏白色短腺毛和黄色开展短腺毛。
  147. 子房被短柔毛
  209d. 腺毛翠雀花 D. grandiflorum var. glandulosum
  147. 心皮无毛
  209e. 光果翠雀花 D. grandiflorum var. leiocarpum
  145. 茎和花梗只被柔毛,无腺毛。
   148. 花序轴和花梗近无毛或有极稀疏毛。
  149. 茎无毛或下部有稀疏硬毛;萼片与距近等长
  207. 秋翠雀花D. autumnale
  149. 茎被极稀疏开展短柔毛
  206. 光序翠雀花 D. kamaonense
   150. 萼距比萼片稍短,长12~16 mm
  206a. 模式变种var. kamaonense
   150. 萼距比萼片长,长18~25 mm
  206b. 展毛光序翠雀花 var. glabrescens
   148. 花序轴和花梗密被柔毛
  209. 翠雀 D. grandiflorum
  151. 茎和花梗只密被贴伏短柔毛。
   152. 子房密被短柔毛
  209a. 模式变种var. grandiflorum
   152. 心皮无毛
  209c. 安泽翠雀花 var. deinocarpum
  151. 茎和花梗密被贴伏短柔毛,此外还有开展长柔毛
  209b. 疏毛翠雀花 var. pilosum
  123. 花序伞房状,或花单朵顶生。
  153. 心皮5。
   154. 萼片比距短,下萼片不呈船形,不具角状突起;退化雄蕊瓣片被黄色髯毛。
  155. 萼距近筒形。
   156. 茎只被白色短柔毛,无黄色腺毛
  212. 宽距翠雀花 D. beesianum
  157. 叶的中央全裂片和侧全裂片的2回裂片均有小裂片。    158. 叶的末回裂片宽2~3.5 mm
  212a. 模式变种var. beesianum
   158. 叶的末回裂片宽3~7 mm
  212b. 粗裂宽距翠雀花var. latisectum
  157. 叶的中央全裂片和侧全裂片的2回裂片均无小裂片
  212c. 辐裂宽距翠雀花var. radiatifolium
   156. 茎被白色短柔毛和黄色短腺毛
  213d. 粗距蓝翠雀花D. caeruleum var. crassicalcaratum
  155. 萼距钻形。
   159. 叶的末回裂片宽1.5~2.5(~4) mm;萼距基部粗3 mm;退化雄蕊瓣片宽倒卵形或近圆形,顶端全缘或微凹
  213. 蓝翠雀花 D. caeruleum
  160. 茎和叶柄被贴伏短柔毛。
   161. 叶片宽达5 cm
  213a. 模式变种 var. caeruleum
   161. 叶片宽达12 cm
  213c. 大叶蓝翠雀花 var. majus
  160. 茎和叶柄被开展短柔毛
  213b. 钝裂蓝翠雀花 var. obtusilobum
   159. 叶的末回裂片宽0.5~1.5 mm; 萼距基部粗2 mm; 退化雄蕊片斜卵形, 2半裂
  215. 细距翠雀花 D. lagarocentrum
   154. 萼片与距近等长; 下萼片近顶端呈船形, 有时具小角状突起; 退化雄蕊瓣片被白色髯毛
  214. 永登翠雀花 D. yongdengense
  153. 心皮3。
   162. 花梗被白色短柔毛和黄色短腺毛。
  163. 花梗全部被白色短柔毛和黄色短腺毛;退化雄蕊瓣片宽倒卵形。
   164. 草本高30~80 cm; 茎被贴伏短柔毛; 伞房花序有3~12花; 萼距长20~25 mm
  217. 康定翠雀花 D. tatsienense
  165. 叶末回裂片宽1.5~2.5(~4) mm;萼片内面无毛
  217a. 模式变种 var. tatsienense
  165. 叶末回裂片宽0.5~1.5 mm;萼片内面有短毛
  217b. 班玛翠雀花 var. chinghaiense
   164. 草本高10~16.5 cm; 茎被开展短柔毛; 伞房花序有(1~)2(~3)花; 萼距长33~38 mm
  218. 矮翠雀花 D. pumilum
  163. 花梗只在顶端被白色短柔毛和黄色短腺毛, 其他部分无毛; 退化雄蕊瓣片宽长圆形, 长13 mm
  219. 硕片翠雀花 D. grandilimbum
   162. 花梗无黄色腺毛。
  166. 退化雄蕊的爪狭楔形,比瓣片短,比瓣片窄2倍
  220. 波密翠雀花 D. pomeense
  166. 退化雄蕊的爪细,与萼片近等长,比瓣片窄5~6倍。
   167. 叶基生并茎生。
  168. 叶片分裂程度小, 末回裂片三角形或狭卵形; 种子椭圆体形, 其纵棱不明显, 无翅
  216. 谷地翠雀花 D. davidii
  168. 叶片强度分裂,全裂片细裂,末回裂片披针状条形、条形或狭条形。
   169. 草本高90~100 cm;叶末回裂片狭条形,宽1.5~1.8 mm;伞房花序有2~5花
  221. 拟蓝翠雀花 D. pseudocaeruleum
   169. 草本高达22~40 cm。
  170. 茎和花梗被开展短柔毛; 叶的末回裂片宽3.5 mm; 萼距长25 mm; 伞房花序有2 花
  222. 軟毛翠雀花 D. mollipilum
  170. 茎和花梗被贴伏短柔毛。
   171. 草本高30~40 cm;花组成具2~3花的伞房花序。
  172. 叶片宽达9.5 cm, 侧全裂片3回细裂, 末回条形裂片宽2.5~4 mm; 退化雄蕊瓣片顶端微凹
  209f. 房山翠雀花 D. grandiflorum var. fangshanense
  172. 叶片宽达5 cm,侧全裂片1回深裂,末回条形裂片宽1~2 mm;退化雄蕊瓣片顶端2微裂
  225. 狭裂翠雀花 D. lagarolobum
   171. 草本高12~22 cm;花单朵顶生;退化雄蕊瓣片2半裂
  226. 李恒翠雀花 D. lihengianum
   167. 叶全部基生。
  173. 花葶被短柔毛, 顶端有1(~2)花; 萼片长16 mm, 距长22~24 mm; 退化雄蕊瓣片2浅裂, 被白色髯毛
  223. 花葶翠雀花 D. sinoscaposum
  173. 花葶被硬毛, 顶端有1花; 萼片长23~24 mm, 距长27 mm; 退化雄蕊瓣片2深裂稍超过中部, 被黄色髯毛
  224. 岩生翠雀花 D. saxatile
  122. 种子具横向的条纹或小鳞片;萼距长达25 mm。
  174. 种子近四面体形, 具横向的条纹; 叶片3深裂至近基部处; 总状或近伞房状花序, 具少数花, 轴和花梗无毛
  227. 长距翠雀花 D. tenii   174. 種子倒卵球形, 具横向排列的小鳞片。
   175. 草本高约100 cm; 茎无毛;萼距长15~22 mm; 花序近伞房状, 有2~3花, 花梗无毛; 心皮无毛
  228. 水城翠雀花 D. shuichengense
   175. 草本高约60 cm; 茎被贴伏短柔毛; 花序伞房状, 有2花, 花梗被贴伏短柔毛; 子房密被短柔毛
  229. 多枝翠雀花 D. maximowiczii
  Key to species and varieties
  1. Petals and staminodes yellowish or yellowish-white; sepals blue or purple.
  2. Leaf blades 3-sect; flowers solitary and terminal; sepals longer than spur; petals yellowish-white; staminode limbs yellowish-white, undivided; carpels 5
  159. Delphinium huangzhongense
  2. Leaf blades 3-parted; flowers in racemes; staminode limbs 2-lobed; carpels 3 or 1.
   3. Sepals nearly as long as spur; petals and staminodes yellowish; carpels 3.
  4. Flowers smaller, in compound racemes; sepals 6-7 mm long, with apexes by right angle incurved; carpels glabrous
  160. D. zhengyianum
  4. Flowers larger, in simple racemes; sepals 14 mm long, not incurved; ovaries puberulous
  161. D. neowentsaii
   3. Sepals shorter than spur; petals and staminodes yellowish-white; carpel l, glabrous
  189. D. monogynum
  1. Petals and staminodes concolour with sepals, i. e. blue and purple, rarely white.
  5. Basal leaves absent, or present, then withered by anthesis.
   6. Seed with 3 longitudinal edges and along edges narrow-winged.
  7. Carpels 4-6.
   8. Leaf blades 3-sect; carpels 5
  98. D. kingianum
  9. Stems hairy; leaf segments at apex acute.
   10. Stens densely white-puberulous and yellow-glandular-puberulous.
  11. Ovaries densely puberulous
  98a. var. kingianum
  11. Ovaries only on ventral sutures hairy, elsewhere glabrous
  98b. var. leiocarpum
   10. Stems densely white-puberulous and mixed with a few yellow glandular hairs
  98c. var. eglandulosum
  9. Stems glabrous; leaf segments at apex long acuminate
  98d. var. acuminatissimum
   8. Leaf blades 3-parted; carpels 4-6
  99. D. humilius
  7. Carpels 3.
   12. Leaf blades 3-sect; sepals longer than spur
  100. D. caudatolobum
   12. Leaf blades 3-parted; sepals nearly as long as spur or shorter than it.
  13. Raceme rachis and pedicels white-puberulous and yellow-glandular-puberulous
  103. D. gyalanum
  13. Raceme rachis and pedicels laching yellow glandular hairs.
   14. Stems and pedicels glabrous.
  15. Herbs up to 140 cm tall; sepals 14-15 mm long; staminode limbs 2-lobed   106. D. cheilanthum
   16. Pedicels retrorsely white-puberulous
  106a. var. cheilanthum
   16. Pedicels spreadingly yellow-puberulous
  106b. var. pubescens
  15. Herbs up to 20-30 cm tall; sepals 15-20 mm long.
   17. Leaf primary lateral lobes 2-cleft; sepals 15-20 mm long, spur 18-24 mm long; staminode limbs suborbicular, 2-lobed
  104. D. latilimbum
   17. Leaf primary lateral lobes 2-parted exceeding the middle of lobe; sepals 15-18 mm long, spur 20 mm long; staminode limbs broad-elliptic, at apex emarginate
  107. D. yuchuanii
   14. Stems and pedicels hairy, rarely stems subglabrous (D. hsinganense).
  18. Sepals on both surfaces hairy, nearly as long as spur or slightly longer than it
  101. D. pachycentrum
   19. Sepals oblong
  101a. var. pachycentrum
   19. Sepals lanceolate
  101b. var. lancisepalum
  18. Sepals inside glabrous.
   20. Compound racemes; bracteoles appressed on flower, narrow-lanceolate, 6-11 mm long; sepals 13 mm long; staminode limbs obovate-oblong, white-barbate
  102. D. pachycentroides
   20. Simple racemes; bracteoles remote from flower, subulate, 3-5 mm long; sepals 14-17 mm long; staminode limbs orbicular-ovate, yellow-barbate
  105. D. hsinganense
   6. Seeds densely transversely squamulose or striate.
  21. Leaf blades broad-cordate, undivided or inconspicuously 3-lobulate
  158. D. calthifolium
  21. Leaf blades pentagonal or reniform, 3-lobed, 3-parted or 3-sect.
   22. Leaf blades 3-lobed or 3-parted.
  23. Leaf blades 3-lobed not exceeding the middle of blade.
   24. Leaf blades 3-lobed not to the middle of blade, lobes not imbricate among themselves; inflorescences racemose, 10 cm long; pedicels white-puberulous and yellow-glandular-puberulous
  135. D. sinovitifolium
   24. Leaf blades 3-cleft to the middle of blade, with lobes imbricate among themselves; inflorescences corymbose, 6 cm long; pedicels glabrous
  134. D. maoxianense
  23. Leaf blades 3-parted exceeding the middle of blades.
   25. Inflorescences corymbose.
  26. Stems glabrous.
   27. Sepals abaxially below apex corniculate
  133. D. furcatocornutum
   27. Sepals not corniculate.
  28. Sepal spur spirally curved
  137. D. bonvalotii
  28. Sepal spur slightly recurved
  130. D. eriostylum
  26. Stems hairy.    29. Stems puberulous; leaf blades up to 6 cm broad
  131. D. pingwuense
   29. Stems hispid; leaf blades up to 12 cm broad.
  30. Pedicels with yellow glandular hairs
  130. D. eriostylum
   31. Pedicels only yellow-glandular-puberulous; ovaries sparsely pubescent
  130a. var. eriostylum
   31. Pedicels white-puberulous and yellow-glandular-puberulous; carpels glabrous
  130b. var. hispidum
  30. Pedicels lacking yellow-glandular hairs
  132. D. laxicymosum
   32. Pedicels glabrous; sepal spur 23-27 mm long
  132a. var. laxicymosum
   32. Pedicels puberulous; sepal spur 20 mm long
  132b. var. pilostachyum
   25. Inflorescences racemose.
  33. Sepal spur straight, slightly or by right angle recurved.
   34. Sepals longer than spur; pedicels 5-9 cm long
  108. D. longipedicellatum
   34. Sepals nearly as long as or shorter than spur.
  35. Sepals nearly as long as spur.
   36. Sepal spur upwards spreading, 10-13 mm long
  110. D. orthocentrum
   36. Sepal spur not upwards spreading.
  37. Stems puberulous; sepal spur horizontally spreading, 12-16 mm long.
   38. Pedicels with few yellow glandular hairs; sepals 12-18 mm long, spur 10-16 mm long; ovaries puberulous
  111. D. micropetalum
   38. Pedicels lacking yellow glandular hairs; sepals 12 mm long,spur 9-12 mm long; carpels glabrous
  109. D. orthocentroides
  37. Stems hispid; sepal spur slightly recurved, 20 mm long
  112. D. hirticaule
   39. Pedicels glabrous
  112a. var. hirticaule
   39. Pedicels with yellow glandular hairs
  112b. var. mollipes
  35. Sepals shorter than spur.
   40. Bracteoles contiguous or nearly contiguous to the flower and appressed to it.
  41. Bracteoles narrow-linear
  122. D. umbrosum
   42. Bractaoles 0.5-1.1 mm broad.
  43. Stems up to 110 cm tall, strigillose; bracteoles 0.5-0.6 mm broad
  122a. var. umbrosum
  43. Stems up to 70 cm tall, hispid; bracteoles 0.6-1.1 mm broad
  122b. var. hispidum
   42. Bracteoles 1.5-1.8 mm broad
  122c. var. drepanocentrum
  41. Bracteoles linear-lanceolate.
   44. Stems strigillose; sepals outside puberulous
  120. D. delavayi             45. Pedicels white-strigillose and yellow-glandular-puberulous
  120a. var. delavayi   45. Pedicels only white-strigllose
  120b. var. baoshanense
   44. Stems hispid; sepals outside sparsely hispid
  121. D. pogonanthum
  46. Stamens glabrous
  121a. var. pogonanthum
  46. Stamen filaments at each side on margin with 1-2 hairs
  121b. var. lasiandrum
   40. Bracteoles remote from flower and not appressed to it.
  47. Leaf central bobe inconspicuously 3-lobed or undivided.
   48. Leaf central lobe rhombic, on margin above the middle densely dentate and below the middle entire.
  49. Sepals 9 mm long, spur 17 mm long; stems strigose or glabrous
  113. D. handelianum
  49. Sepals 15-16 mm long, spur 20-21 mm long; stems glabrous
  114. D. honanense
   50. Pedicels white-puberulous and yellow-glandular-puberulous
  114a. var. honanense
   50. Pedicels only white-puberulous
  114b. var. piliferum
   48. Leaf central lobe lanceolate, on margin from above base to apex densely dentate
  115. D. hirtifolium
  47. Leaf central lobe rhombic, near the middle conspicuously 3-lobed, rarely 3-parted.
   51. Leaf central lobe wide rhombic, near midrib 3-parted
  129. D. pseudocyananthum
   51. Leaf central lobe rhombic, 3-lobed.
  52. Stems glabrous.
   53. Pedicels glabrous
  136. D. potaninii
  54. Carpels glabrous; follicles straight, oblong, 1.4-1.7 cm long.
   55. Bracteoles 0.2-0.3 mm broad
  136a. var. potaninii
   55. Bracteoles 1-1.2 mm broad
  136b. var. latibracteolatum
  54. Carpels with sparse hairs; follicles lunated, 0.5-1 cm long
  136c. var. jiufengshanense
   53. Pedicels hairy.
  56. Pedicels densely yellow-glandular-puberulous.
   57. Racemes compound; carpels glabrous.
  58. Herbs 1.5-2 m tall; pedicels white-strigillose and yellow-glandular-puberulous; sepals 11 mm long, spur 18 mm long; staminode limbs 2-parted exceeding the middle of limb
  125. D. polyanthum
  58. Herbs ca. 80 cm tall; pedicels only yellow-glandular-puberulous; sepals 7-10 mm long, spur 11-13 mm long; staminode limbs at apex emarginate
  126. D. zuogongense
   57. Racemes simple; ovaries pubescent
  127. D. tongolense
  59. Petioles glabrous
  127a. var. tongolense
  59. Petioles hirsute
  127b. var. hirsutipes
  56. Pedicels only white-puberulous, lacking yellow glandular hairs; racemes compound; herbs ca. 1 m tall   128. D. altissimum
  52. Stems hairy.
   60. Stems and pedicels white-puberulous and yellow-glandular-puberulous
  123. D. sherriffii
   60. Stems and pedicels lacking yellow glandular hairs.
  61. Stems hispid; staminode limbs yellow-barbate.
   62. Racemes simple; sepals outside white-puberulous and yellow-glandular-puberulous; staminode claws slender, ca. 5 times narrower than limbs
  116. D. omeiense
  63. Sepals 12-16 mm long, spur 20-26 mm long
  116a. var. omeiense
  63. Sepals 4-8 mm long, spur 7-18 mm long
  116b. var. micranthum
   62. Racemes compound; sepals outside only with yellow glandular hairs, 10 mm long, spur 20 mm long; staminode claws linear, slightly narrower than limbs
  119. D. platyonychium
  61. Stems pubescent.
  64. Stems spreading-pubescent; racemes compound; sepal spur 23 mm long
  124. D. yanyuanense
  64. Stems retrorsely puberulous; racemes simple; sepal spur 17-18 mm long.
   65. Leaf blades 3-parted to 2-4 mm above base; upper sepal narrow-ovate, 13 × 6 mm; staminode limbs yellow-barbate
  117. D. pubescens
   65. Leaf blades 3-parted to 6-7 mm above base; upper sepal broud-ovate, 10 × 7 mm; staminode limbs white-barbate
  118. D. zhouchengense
  33. Sepal spur by U-form or spirally recurved.
   66. Stems glabrous.
  67. Pedicels glabrous
  136. D. potaninii
  67. Pedicels hairy.
   68. Pedicels only yellow-glandular-puberulous; leaf blades up to 16 cm broad, 3-parted slightly exceeding the middle of blade; bracteoles linear
  141. D. wenchuanense
   68. Pedicels white-puberulous and yellow-glandular-puberulous; leaf blades up to 12 cm broad, 3-parted to near the base of blade; bracteoles filiform.
  69. Bracteoles 4.5-7 mm long; sepal spur 12-21 mm long
  140. D. pseudotongolense
  69. Bracteoles 1.5-4 mm long; sepal spur 18-24 mm long
  138. D. bulleyanum
   70. Ovaries spasely pubescent
  138a. var. bulleyanum
   70. Carpels glabrous
  138b. var. leiogynum
   66. Stems hairy.
  71. Carpels glabrous
  139. D. gonggaense
  71. Ovaries hairy.
   72. Stems hispid; leaf blades up to 10(-14) cm broad; sepals 15-18 mm long
  142. D. spirocentrum
   72. Stems spreading-puberulous; leaf blade up to 5.4 cm broad; sepals 12-13 mm long
  143. D. angustirhombicum    22. Leaf blades 3-sect.
  73. Leaf segments dissected, with ultimate lobes linear-lanceolate
  157. D. aemulans
  73. Leaf segments not dissected, with ultimate lobes ovate or triangular.
   74. Sepal spur less than 15 mm long.
  75. Stems,pedicels and carpels glabrous
  144. D. sparsiflorum
  75. Stems and pedicels hairy.
   76. Herbs ca. 100 cm tall; racemes compound; sepal spur 8-9 mm long
  145. D. angustipaniculatum
   76. Herbs 60-80 cm tall; racemes simple; sepal spur 12-15 mm long.
  77. Leaf central lobe rhombic, at the middle 3-lobed or subpinnately divided; ovaries pubescent
  146. D. conaense
  77. Leaf central broad-rhombic, below the middle 3-parted; carpels glabrous
  147. D. wangii
   74. Sepal spur 20-28 mm long.
  78. Stems glabrous.
   79. Sepals not corniculate.
  80. Pedicels glabrous.
   81. Herbs up to 110 (-150) cm tall; leaf blades up to 20 cm broad; racemes compound; bracteoles 2.5-3.5 mm long; sepal spur straight or recurved
  151. D. giraldii          81. Herbs up to 70 cm tall; leaf blades up to 8 cm broad; racemes simple; bracteoles 4-5 mm long; sepal spur spirally recurved
  152. D. glabricaule
  80. Pedicels hairy.
   82. Pedicels white-puberulous and yellow-glandular-puberulous; racemes simple
  148. D. muliense
  83. Bracteoles lanceolate or lanceolate-linear, 8-13 × 1.2-2 mm
  148a. var. muliense
  83. Bracteoles subfiliform, 3.5-4 × 0.2 mm
  148b. var. minutibracteolatum
   82. Pedicels only white-puberulous; racemes compound
  153. D. sutchuenense
   79. Sepals outside below apex corniculate.
  84. Pedicels white-puberulous; sepal horn-like projections minute, inconspicuous; spur horizontally spreading or slightly recurved
  153. D. sutchuenense
  84. Pedicels yellow-glandular-puberulous; sepals with conspicuous horn-like projections 2.5-3.5 mm long, spur by U-form or spirally recurved
  154. D. campylocentrum
  78. Stems hairy.
   85. Pedicels glabrous.
  86. Leaf blades abaxially glabrous; sepals glabrous, spur 28 mm long
  156. D. dazangense
  86. Leaf blades abaxially hairy; sepals hairy, spur 20 mm long.
   87. Stems below yellow-hispid; sepals inside glabrous
  149. D. rangtangense
   87. Stems below white-hispid; sepals inside with sparse hairs   150b. D. pseudocampylocentrum var. glabripes
   85. Pedicels hairy.
  88. Herbs ca. 50 cm tall; pedicels sparsely yellow-glandular-puberulous; bracteoles remote from flower; sepal spur 17-22 mm long; staminode limbs yellow-barbate; ovaries with sparse hairs
  150a. D. pseudocampylocentrum var. pseudocampylocentrum
  88. Herbs 50-90 (-100) cm tall; pedicels densely white-puberulous; bracteoles contiguous to flower; sepal spur 22-25 mm long; staminode limbs not barbate; carpels glabrous
  155. D. trifoliolatum
  5. Basal leaves well developed, rarely absent or present but withered by anthesis.
   89. Leaf blades 3-lobed or 3-parted.
  90. Seeds subtetrahedral, along edges narrow-winged.
   91. Sepals outside below apex corniculate.
  92. Leaf blades 3-parted slightly exceeding the middle of blade,with ultimate lobes ovate or broad-triangular, rarely narrow-lanceolate or lanceolate-linear.
   93. Leaf central lobe thrice divided
  166. D. taliense
  94. Leaf ultimate lobes ovate or triangular.
   95. Sepal spur at base 2-3 mm across.
  96. Sepals outside puberulous.
   97. Stem glabrous.
  98. Carpels glabrous
  166a. var. taliense
  98. Ovaries puberulous
  166b. var. glabrum
   97. Stems below densely puberulous
  166c.var. pubicaule
  96. Sepals outside hirsute
  166d. var. hirsutum
   95. Sepal spur at base 4-5 mm across
  166e. var. platycentrum
  94. Leaf ultimate lobes narrow-lanceolate or lanceolate-linear; sepal spur 2 times longer than sepals
  166f. var. dolichocentrum
   93. Leaf central lobe once or twice divided
  167. D. ceratophorum
  99. Sepals with horn-like projections 1-2 mm long.
   100. Leaf blades up to 10 cm broad, ultimate lobes ovate or triangular.
  101. Sepal outside sparsely strigillose (hairs ca. 0.4 mm long), with horn-like projections 1.5-2 mm long
  167a. var. ceratophorum
  101. Sepals outside hirsute (hairs ca. 2 mm long), with horn-like projections 1-1.5 mm long
  167c. var. hirsutum
   100. Leaf blades 8-19 cm broad, ultimate lobes linear-lanceolate; sepal horn-like projections 1.5-2 mm long
  167d. var. robustum
  99. Sepal horn-like projections less than 1 mm long
  167b. var. brevicorniculatum
  92. Leaf blades 3-parted to near the base of blade.
   102. Pedicels above yellow-glandular-puberulous and white-hispid; sepals 10-12 mm long; staminode limbs white-barbate   169. D. brevisepalum
   102. Pedicels glabrous; sepals 14-22 mm long; staminode limbs yellow-barbate.
  103. Stems glabrous; sepals 18-22 mm long, horn-like projections 1 mm long, spur 24 mm long
  168. D. ceratophoroides
  103. Stems below strigose; sepals 14-16 mm long, horn-like projections 1-1.5 mm long, spur 27-28 mm long
  170. D. dolichocentroides
   91. Sepals not corniculate.
  104. Leaf blades 3-cleft or 3-parted slightly exceeding the middle of blade.
   105. Pedicels glabrous
  162. D. hamatum
   105. Pedicels white-puberulous and yellow-glandular-puberulous.
  106. Stems near apex yellow-glandular-puberulous; sepals 9 mm long, spur 14-17 mm long; carpels glabrous
  163. D. latirhombicum
  106. Stems glabrous; sepals 15-19 mm long; ovaries pubescent
  165. D. ninglangshanicum
  104. Leaf blades 3-parted to near the base of blade.
   107. Sepals 13-19 mm long.
  108. Pedicels lacking yellow glandular hairs.
   109. Pedicels above 2-bracteolate.
  110. Racemes 2-6.5(-16) cm long; ovaries densely pubescent.
   111. Racemes 16 cm long; pedicels 3.5-10 cm long
  174. D. yanwaense
   111. Racemes 2-6.5 cm long; pedicels 1.4-3.2 cm long.
  112. Sepal spur 25-27 mm long; staminode limbs 2-cleft, white-barbate
  172. D. weixiense
  112. Sepal spur 18 mm long; staminode limbs at apex emarginate, yellow-barbate
  186. D. lankongense
  110. Racemes 10-26 cm long; ovaries only on ventral sutures hairy, elsewher glabrous
  187. D. pycnocentrum
   109. Pedicels on apex 2-bracteolate (bracteoles linear)
  164. D. umbrosoides
  108. Pedicels yellow-glandular-puberulous and white-puberulous; sepal spur 16-18 mm long.
   113. Stem glabrous; stamen filaments hairy.
  113′. Racemes 22 cm long; staminode limbs obcordate, 7 mm broad; anthers glabrous
  182. D. obcordatilimbum
  113′. Racemes 12-15 cm long; staminode limbs oblong, 3 mm broad; anthers abaxially hair
  179. D. lasiantherum
   113. Stems hary; filaments glabrous.
  113″. Stems puberulous, 1-leaved; basal leaves ca. 7; staminode limbs 2-parted slightly exceedling the middle of limb, white-barbate; ovaries puberulous
  171. D. incisolobulatum
  113″. Stems strigose, several-leaved; basal leaves withered by anthesis; staminode limbs undivided or at apex 2-lobulate, yellow-barbate; carpels glabrous   173. D. kantzeense
   107. Sepals 20-30 mm long.
  114. Leaf sheaths well developed, 8-15 mm broad
  180. D. coleopodum
  114. Leaf sheaths narrow or absent.
   115. Pedicels glabrous.
  116. Stems puberulous; sepals 9-14 mm long, spur 17-24 mm long; staminode limbs 2-cleft
  178. D. yunnanense
  116. Stems glabrous; sepals 18 mm long, spur 26-30 mm long; staminode limbs undivided
  185. D. liangshanense
   115. Pedicels hairy.
  117. Pedicels with yellow glamdular hairs.
   118. Stems hispid; leaf blades on both surfaces sparsely strigose; sepal spur 17-26 mm long
  175. D. yongningense
   118. Stems puberulous; leaf blades glabrous; sepal spur 22 mm long
  176. D. pergameneum
  117. Pedicels lacking yellow glandular hairs.
   119. Herbs 3-8 cm tall; pedicels white-puberulous; sepals 15-16 mm long, spur 25 mm long
  181. D. hui
  120. Leaf blade with central lobe 3-dentate; stem apex 1-2-flowered
  181a. var. hui
  120. Leaf blade with central lobe 3-5-dentate; stem apex 1-5(-10)-flowered
  181b. var. lobulibundum
   119. Herbs 11-18 cm tall; pedicels yellow-puberulous; sepal spur 28-29 mm long.
  121. Stems below sparsely spreading-puberulous; bracteoles 5 mm long; staminode limbs at apex rounded and entire
  183. D. pseudohamatum
  121. Stems retrorse-puberulous; bracteoles (5-)7.5-9 mm long; staminode limbs at apex emarginate or 2-lobulate
  184. D. yulongshanicum
  90. Seeds densely transversely squamulose; leaf blades 3-parted slightly exceeding the middle of blade; sepals 13-15 mm long, spur 20-22 mm long
  188. D. hueizeense
   89. Leaf blades 3-sect.
  122. Seeds subtetrahedral, along longitudinal edges narrow-winged.
   123. Inflorescences racemose.
  124. Upper sepal eymbiform.
   125. Leaf blades at base deep-cordate, ultimate lobes triangular or narrow-lanceolate; sepal spur shorter than sepals, tubular-subulate, 11-18 mm long
  190. D. likiangense
   125. Leaf blades at base subtruncate, ultimate lobes linear; sepal spur longer than sepals, subtubular, 21-27(-30) mm long
  191. D. souliei
  124. Upper sepal more or less flat, not cymbiform.
   126. Leaf ultimate lobes ovate, triangular or narrow-lanceolate.
  127. Sepals longer than spur or nearly as long as it.
   128. Stems and pedicels puberulous; sepals longer than spur, 17-23 mm long; spur 12-16 mm long   192. D. medogense
   128. Stems and pedicels glabrous; sepals and spur 10-12 mm long
  193. D. yuanum
  127. Sepals shorter than spur.
   129. Stems densely spreading-puberulous
  210. D. weihaiense
   129. Stems appressed-puberulous, hispid or glabrous.
  130. Pedicels yellow-glandular-puberulous and white-puberulous
  204b. D. thibeticum var. laceratilobum
  130. Pedicels lacking yellow glandular hairs.
   131. Pedicels glabrous; leaves all basal
  199. D. breviscaposum
   131. Pedicels hairy.
  132. Stems and carpels glabrous; sepal spur 15 mm long
  194. D. filibracteolatum
  132. Stems and carpels hairy.
   133. Staminode limbs undivided.
  134. Sepal spur 24-28 mm long; staminode limbs at apex erose
  201. D. nangchienense
  134. Sepal spur 16-24 mm long; staminode limbs at apex emarginate.
   135. Bracteoles narrow-oblong, 11-18 mm long; sepal spur up to 19 mm long
  195. D. weiningense
   135. Bracteoles narrow-linear or subulate, 2.5-6 mm long; sepal spur up to 24 mm long
  200. D. majus
   133. Staminode limbs 2-lobed.
  136. Leaves all basal, central segment subcuneate, once divided, near apex 3-lobed or 3-dentate
  198. D. chayuense
  136. Leaves basal and cauline, central segment thrice divided.
  137. Stems retrorse-puberulous; leaf central segment cuneate-rhombic; bracteoles oblong, 9-12 mm long; sepals 15-18 mm long, spur 14-20 mm long
  196. D. taipaicum
  137. Stems spreading-hispid; leaf central segment rhombic; bracteoles linear, 7-8 mm long; sepals 10-14 mm long, spur 20-24 mm long
  197. D. pseudothibeticum
   126. Leaf segments dissected, with ultimate lobes lanceolate-linear, linear or narrow-linear.
  138. Carpels 5.
   139. Raceme rachis and pedicels retrorse-puberulous, lacking yellow glandular hairs; sepals 9 mm long, spur 12 mm long; staminode limbs at apex emarginate; carpels glabrous
  202. D. sinopentagynum
  139. Raceme rachis and pedicels with yellow glandular hairs; sepals 15 mm long, spur 20-24 mm long; staminode limbs at apex 2-lobulate; ovaries pubescent
  203. D. pseudomosoynense
  140. Raceme rachis and pedicels white-puberulous and yellow-glandular-puberulous; upper sepal outside below apex with a small horn-like projection, spur 20 mm long; petals at apex ciliate
  203a. var. pseudomosoynense   140. Raceme rachis and pedicels only yellow-glandular -puberulous, lacking white hairs; upper sepal lacking horn-like projection, spur 21-24 mm long; petals glabrous
  203b. var. subglabrum
  138. Carpels 3.
   141. Staminode limbs oblong or obovate-oblong.
  142. Raceme rachis and pedicels glabrous.
   143. Leaf ultimate lobes 6 mm broad; racemes 25-45 cm long; sepal spur 13-15 mm long; staminode limbs 4.6 mm long, 2-cleft
  177. D. pseudoyunnanense
   143. Leaf ultimate lobes 1.6-3 mm broad; racemes 10-20 cm long; sepal spur 25-28 mm long; staminode limbs 8 mm long, 2-lobulate
  205. D. leiostachyum
  142. Raceme rachis and pedicels hairy.
   144. Raceme rachis and pedicels only white-puberulous; bracteoles remote from flower
  208. D. mosoynense
   144. Raceme rachis and pedicels white-puberalous and yellow-glandular-puberulous; bracteoles contiguous to flower
  204a. D. thibeticum var. thibeticum
   141. Staminode limbs broad-obovate or suborbicular.
  145. Stems and pedicels puberulous and glandular-puberulous.
   146. Stems and pedicels densely spreading-white-puberulous and white-glandular- pube-rulous
  211. D. chefoense
   146. Stems and pedicels densely appressed-white-puberulous and spreading-yellow-puberulous.
  147. Ovaries puberulous
  209d. D. grandiflorum var. glandulosum
  147. Carpels glabrous
  209e. D. grandiflorum var. leiocarpum
  145. Stems and pedicels only puberulous, not glandular-puberulous.
   148. Raceme rachis and pedicels subglabrous or with very sparse hairs.
  149. Stems glabrous or below sparsely hispid; sepals nearly as long as spur
  207. D. autumnale
  149. Stems with very sparse spreading hairs
  206. D. kamaonense
   150. Sepal spur slightly shorter than sepals, 12-16 mm long
  206a. var. kamaonense
   150. Sepal spur longer than sepals, 18-25 mm long
  206b. var. glabrescens
   148. Raceme rachis and pedicels with dense hairs
  209. D. grandiflorum
  151. Stems and pedicels only densely puberulous.
   152. Ovaries densely appressed-puberulous
  209a. var. grandiflorum
   152. Carpels glabrous
  209c. var. deinocarpum
  151. Stems and pedicels densely appressed-puberulous, and with some spreading longer hairs
  209b. var. pilosum
   123. Inflorescences corymbose, or flowers solitary and terminal.   153. Carpels 5.
   154. Sepals shorter than spur, lower sepal not, not corniculate; staminode limbs yellow-barbate.
  155. Sepal spur subtubular.
   156. Stems only white-puberulous, lacking yellow glandular hairs
  212. D. beesianum
  157. Leaf central segment and secondary lobes of lateral segments all with lobules.
   158. Leaf ultimate lobes 2-3.5 mm broad
  212a. var. beesianum
   158. Leaf ultimate lobes 3-7 mm broad
  212b. var. latisectum
  157. Leaf central segment and secondary lobes of lateral segments all lacking lobules
  212c. var. radiatifolium
   156. Stems white-puberulous and yellow-glandular-puberulous
  213d. D. caeruleum var. crassicalcaratum
  155. Sepal spur subulate.
   159. Leaf ultimate lobes 1.5-2.5(-4) mm broad; sepal spur at base 3 mm across; staminode limbs broad-obovate or suborbicular, at apex entire or emarginate
  213. D. caeruleum
  160. Stems and petioles appressed-puberulous.
   161. Leaf blades up to 5 cm broad
  213a. var. caeruleum
   161. Leaf blades up to 12 cm broad
  213c. var. majus
  160. Stems and petioles spreading-puberulous
  213b. var. obtusilobum
   159. Leaf ultimate lobes 0.5-1.5 mm broad; sepal spur at base 2 mm across; staminode limbs white-barbate
  215. D. lagarocentrum
   154. Sepals nearly as long as spur; lower sepals near apex cymbiform, sometimes minutely 1-corniculate; staminode limbs white-barbate
  214. D. yongdengense
  153. Carpels 3.
   162. Pedicels white-puberulous and yellow-glandular-puberulous.
  163. Pedicels totally white-puberulous and yellow-glandular-puberulous; staminode limbs broad-obovate.
   164. Herbs 30-80 cm tall; stems appressed-puberulous; corymbs 3-12-flowered; sepal spur 20-25 mm long
  217. D. tatsienense
  165. Leaf ultimate lobes 1.5-2.5(-4) mm broad; sepals inside glabrous
  217a. var. tatsienense
  165. Leaf ultimate lobes 0.5-1.5 mm broad; sepals inside with short hairs
  217b. var. chinghaiense
   164. Herbs 10-16.5 cm tall; stems spreading-puberulous; corymbs (1-)2(-3)-flowered; sepal spur 33-38 mm long
  218. D. pumilum
  163. Pedicels only on apex white-puberulous and yellow-glandular-puberulous, elsewhere glabrous; staminode limbs broad-oblong, 13 mm long
  219. D. grandilimbum
   162. Pedicels lacking yellow glandular hairs.   166. Staminode claws narrow-cuneate, shorter than limbs and two times narrower than limbs
  220. D. pomeense
  166. Staminode claws slender, nearly as long as limbs and 5-6 times narrower than them.
   167. Leaves basal and cauline.
  168. Leaf blades weakly divided, with ultimate lobes triangular or narrow ovate; seeds ellipsoidal, with inconspicuous longitudinal edges and not winged
  216. D. davidii
  168. Leaf blades strongly divided, with segments dissected and ultimate lobes lanceolate- linear, linear or narrow-linear.
   169. Herbs 90-100 cm tall; leaf ultimate lobes narrow-linear, 1.5-1.8 mm broad; corymbs 2-5-flowered
  221. D. pseudocaeruleum
   169. Herbs up to 22-40 cm tall.
  170. Stems and pedicels spreading-puberulous; leaf ultimate lobes 3.5 mm broad; sepal spur 25 mm long; corymbs 2-flowered
  222. D. mollipilum
  170. Stems and pedicels appressed-puberulous.
   171. Herbs 30-40 cm tall; flowers in 2-3-flowered corymbs.
  172. Leaf blades up to 9.5 cm broad, lateral segments thrice dissected, with ultimate lobes 2.5-4 mm broad; staminode limbs at apex emarginate
  209f. D. grandiflorum var. fangshanense
  172. Leaf blades up to 5 cm broad, lateral segments once parted, with ultimate lobes 1-2 mm broad; staminode limbs at apex 2-lobulated
  225. D. lagarolobum
   171. Herbs 12-22 cm tall; flowers solitary and terminal; staminode limbs 2-cleft
  226. D. lihengianum
   167. Leaves all basal.
  173. Scapes puberulous, on apex 1(-2)-flowered; sepals 16 mm long, spur 22-24 mm long; staminode limbs 2-lobed
  223. D. sinoscaposum
  173. Scapes hispid, on apex 1-flowered; sepals 23-24 mm long, spur 27 mm long; staminode limbs 2-parted slightly exceeding the middle of limb
  224. D. saxatile
  122. Seeds transversely striate or squamulose.
   174. Seed subtetrahedral, transversely striate; leaf blades 3-parted to near base of blade; racemes subcorymbose, few-flowered
  227. D. tenii
   174. Seeds obovoid, transversely squamulose.
  175. Herbs ca. 100 cm tall; stems glabrous; inflorescences subcorymbose, 2-3-flowered, pedicels glabrous; carpels glabrous
  228. D. shuichengense
  175. Herbs ca. 60 cm tall, appressed-puberulous; inflorescences corymbose 2-flowered, pedicels appressed-puberuous; ovaries puberulous
  229. D. maxinowiczii
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