Fluctuations in food availability occur in many ecosystems and pose challenges to the organisms that inhabit them(Berg et al.2017).Because lizards have significantly lower field metabolic rates than endotherms,they are often considered “low energy specialists” that can cope with food shortages(Brown and Griffin 2005).However,active reptiles that maintain high body temperatures during periods of low prey availability can still incur significant energetic costs be-cause metabolic rates scale with body temperature.