中国银莲花属新分类

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  摘 要:  該文对中国毛茛科银莲花属(Anemone L.)进行了分类学研究,共收载86种,可将这些种划分为10组:其中5组,即草玉梅组、西南银莲花组、钝裂银莲花组、短柱银莲花组和直果银莲花组均是该属具中等演化水平的分类群;另外5组,即银莲花组、鹅掌草组、卵叶银莲花组、二岐银莲花组和打破碗花花组则均是具高度演化水平的分类群。在这10组中,钝裂银莲花组与银莲花组有相近的亲缘关系,短柱银莲花组与鹅掌草组有相近的亲缘关系,而其余6组之间的亲缘关系情况,目前尚未了解。该文写出了区别所有属下分类群的检索表、所有分类群的文献引证和地理分布,并对每一种均指出了研究的重要标本,对多数种均提供插图。描述了2新组(短柱银莲花组和直果银莲花组)、4新系(鹅掌草组的鹤峰银莲花系和加查银莲花系,以及直果银莲花组的兴义银莲花系和直果银莲花系)、13新种(九龙银莲花、柏枝山银莲花、狭萼银莲花、道孚银莲花、泸定银莲花、波密银莲花、嘎隆拉山银莲花、三小叶银莲花、折多山银莲花、宝兴银莲花、贡嘎银莲花、福贡银莲花和盘蕊银莲花)、1新亚种(长柄钝裂银莲花)以及4新变种(紫萼折多山银莲花、矮展毛银莲花、疏毛川西银莲花和泸水银莲花)。此外,还将变种条叶银莲花提升为种级分类群;将涪陵银莲花降级作为川西银莲花的一个变种;由于Anemone rupestris Hook. f. & Thoms. 为一不合法的晚出异物同名,遂拟出一新名(Anemone neorupestris W. T. Wang);小花草玉梅在过去多年间被处理为草玉梅的一个变种,在该文中其原来种级群的地位得到恢复;1939年,H. Handel-Mazzetti将变种云南银莲花归并于展毛银莲花,将稀果银莲花归并于滇川银莲花,对此二归并,该文赞同并采用。
  关键词: 毛茛科,银莲花属, 分类, 中国
  中图分类号:  Q949
  文献标识码:  A
  文章编号:  1000-3142(2021)增刊1-0001-118
  A new classification of Anemone (Ranunculaceae) of China
  WANG Wen-tsai
  ( State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences, Beijing 100093, China )
  Abstract:  A taxonomic study of the genus Anemone (Ranunculaceae) of China has been conducted. Eighty-six species are recognized and classified into ten sections. Of them, five sections, Rivularidium, Anemonanthea, Himalayicae, Brachycladus and Pleurocarpus, are groups with medium-evolutionary level, and the another five sections, Omalocarpus, Stolonifera,Begoniifolia, Anemonidium and Anemone, are groups with high-evolutionary level. In these ten sections,Sect. Himalayicae is closely related to Sect. Omalocarpus, Sect. Brachycladus is closely related to Sect. Stolonifera, and the relationships among the other six sections are not yet clear  now. A key to sections, series, species and infraspecific taxa, and literature citations and geographic distributions for all taxa are presented. Records of important specimens studied for each species and illustrations for most species are provided. Two sections (Sects. Brachycladus and Pleurocarpus), four series (Ser. Hofengenses and Ser. Jiachaenses under Sect. Stolonifera and Ser. Xingyienses and Ser. Orthocarpae under Sect. Pleurocarpus), thirteen  species (A. jiulongensis, A. baizhishanica, A. stenosepala, A. daofuensis, A. ludingensis, A. bomiensis, A. galonglashanica, A. trifoliolata, A. zheduoshanica, A. baoxingensis, A. gonggaensis, A. fugongensis and A. discoideogyna), one subspecies (A. obtusiloba ssp. dolichopoda), and four varieties (A. zheduoshanica var. atropurpurea, A. prattii var. sparsipilifera,  A. fugongensis var. lushuiensis and A. demissa var. humilis) are described as new. Besides, A. obtusiloba ssp. trullifolia var. linearis Bruhl is elevated to specific rank; A. fulingensis is relegated to a variety under A. prattii; due to that A. rupestris Hook. f. & Thoms. is an illegitimate later homonym, its specific epithet ‘rupestris’ is changed as ‘neorupestris’ in this paper; A. barbulata, long relegated to a variety under A. rivularis, is restored to its previous specific rank; and the reduction of A. demissa var. yunnanensis Franch. to var. demissa, and the reduction of A. oligocarpa Pei to A. delavayi Franch, by H. Handel-Mazzetti in 1939 are accepted in this paper.   Key words: Ranunculaceae, Anemone, classification, China
  《中國植物志》的中国银莲花属(Anemone L.)(Wang, 1980)收载的53种被划分为8组,即西南银莲花组(Sect. Anemonanthea)(12种)、鹅掌草组(Sect. Stolonifera)(8种)、草玉梅组(Sect. Rivularidium)(2种)、卵叶银莲花组(Sect. Begoniifolia)(4种)、打破碗花花组(Sect. Anemone)(8种)、钝裂银莲花组(Sect. Himalayicae)(9种)、银莲花组(Sect. Omalocarpus)(9种)和二歧银莲花组(Sect. Anemonidium)(1种)。当时,我只认识到具扁平瘦果的2组银莲花组(Sect.  Omalocarpus)和二歧银莲花组(Sect. Anemonidium)是此属的进化群,将此2组放在分类系统中的最后位置。后来,我看到日本植物学家M. Tamura(田村道夫)教授的此属分类系统将心皮和瘦果数目多、密被长绵毛的打破碗花花组(Sect. Anemone)放在其系统的最后位置(Tamura, 1995),我赞同他的处理。此外,在研究了此属植物花的构造后,我还认识到鹅掌草组(Sect. Stolonifera)[心皮花柱极短或不存在,柱头发育,呈球形、卵球形等形状,花粉具4~10沟(Xi & Chang, 1964)]和卵叶银莲花组(Sect. Begoniifolia)[心皮具短花柱和相当发育的椭圆体形柱头,瘦果具4条纵肋,卵叶银莲花(A. begoniifolia)的花粉具13~18个散孔(Xi & Chang, 1964; Zhang et al., 2015)] 2组均具明显进化特征,也是此属的进化群。 Hoot et al. (1994) 一方面认为卵叶银莲花(A. begoniifolia)和 A. glaucifolia应放在分布于非洲的A. caffra群中;另一方面又认为在瘦果形态上则与草玉梅(A. rivularis)群极为相似。而Tamura(1995)却将卵叶银莲花组(Sect. Begoniifolia)放在Subgen. Omalocarpus中的钝裂银莲花组(Sect. Himalayica)和银莲花组(Sect. Omalocarpus)2组之前。其他3组,即草玉梅组(Sect. Rivularidium)、西南银莲花组(Sect. Anemonanthea)、钝裂银莲花组(Sect. Himalayicae)则均比上述5组原始,它们既有一些原始特征,又有一些进化特征。Tamura (1995)认为草玉梅组(Sect. Rivularidium)无特化特征,是银莲花属的原始群,并且将此组放在其银莲花属分类系统的首位;Tamura的观点得到Ehrendorfer & Samuel (2001)分子系统学研究的赞同。此组的总苞苞片具柄,心皮的柱头不明显,这些的确均是原始特征,但此组的心皮的花柱较长,并钩状弯曲,到果期变硬,可能是适应动物传播果实的结果。此外,小花草玉梅(Anemone barbulata)和草玉梅(A. rivularis)的花粉均具12条散沟,是高度进化现象(Xi & Chang, 1964)。另一组,西南银莲花组(Sect. Anemnanthea),此组的总苞苞片具柄,心皮的花柱通常比子房稍短,直或稍弯曲,这些都是原始特征,在此组多数种中,心皮具明显柱头,则是进化的特征。关于钝裂银莲花组(Sect. Himalayicae),此组雄蕊花丝条形,心皮的花柱比子房稍短、直,柱头通常不明显,这些均是原始特征,但此组的总苞苞片的柄完全消失,显示出高度的退化现象,其染色体基数x=7也是一次生现象。此外,关于短柱银莲花(A. brevistyla),此种的心皮构造与上述钝裂银花莲组(Sect. Himalayicae)的心皮十分相似,花柱直,比子房稍短,柱头不明显,其植株体态(habit)和雄蕊花丝丝形以及花粉具6~7条沟的情况,与鹅掌草组(Sect. Stolonifera)相同。在《中国植物志》中,我将此种置于鹅掌草组(Sect. Stolonifera)中作为第一个种,认为此种是此组的原始种。但鹅掌草组(Sect. Stolonifera)植物的心皮花柱极短或不存在,柱头极为发育,呈球形、卵球形等形状,这种心皮构造与短柱银莲花(A.brevistyla)的心皮构造大相径庭。最近,我发现产自四川贡嘎山的一新种,即贡嘎银莲花(A. gonggaensis),其心皮构造与短柱银莲花(A. brevistyla)的心皮构造近似,区别在于其心皮花柱顶端具一小但明显的柱头。我根据此2种建立一新组,名为短柱银莲花组(Sect. Brachycladus),此新组的花构造较原始,但短柱银莲花(A. brevistyla)的花粉具6~7沟,则是进化的特征 [在银莲花属中,多数种均具三沟花粉(Xi & Chang, 1964)]。此外,提到直果银莲花(A. orthocarpa)(产自贵州兴义),Handel-Mazzetti 在1939年发表此新种时将此种与草玉梅(A. rivularis)作比较,但此种心皮的钻形花柱不呈钩状弯曲,其瘦果具4条肋,花粉具6沟,因此与草玉梅(A. rivularis)有明显区别。在《中国植物志》中,我将此种放在心皮无毛,瘦果也具4条肋的卵叶银莲花组(Sect. Begoniifolia)中,如前所述,卵叶银莲花组(Sect. Begoniifolia)的心皮的花柱极短或不存在,柱头较发育,呈椭圆体形,这与直果银莲花(A. orthocarpa)的心皮构造(花柱较长,呈钻形,柱头不明显)大不相同。最近,我发现特产贵州、兴义的兴义银莲花(A. xingyiensis)的心皮具短但明显的花柱,柱头不明显,瘦果具肋,在这些特征方面与直果银莲花(A. orthocarpa)相似,于是根据此2种建立另一新组,名之为直果银莲花组(Sect. Pleurocarpus),此组的心皮形态显示出原始特征,但在具肋瘦果方面则显示出进化特征。目前,中国银莲花属的种数已增至86种,在本文中被划分为10组,其中草玉梅组(Sect. Rivularidium)、西南银莲花组(Sect. Anemonanthea)、钝裂银莲花组(Sect. Himalayicae)、短柱银莲花组(Sect. Brachycladus)和直果银莲花组(Sect. Pleurocarpus)5组既有原始特征,又有进化特征,是具中等演化水平的分类群;其余5组,即银莲花组(Sect. Omalocarpus)、鹅掌草组(Sect. Stolonifera)、卵叶银莲花组(Sect. Begoniifolia)、二岐银莲花组(Sect. Anemonidium)和打破碗花花组(Sect. Anemone)则均是具高度演化水平的分类群。最近,我看到一篇关于银莲花属卵叶银莲花组的分子系统学文章(Zhang et al., 2015),该文将瘦果被绵毛,以野棉花(A. vitifolia)为代表的Subsect. Vitifoliae和以岩生银莲花(A. rupicola)为代表的Subsect. Rupicolae,以及心皮无毛、花柱短、柱头明显、瘦果具4条纵肋的卵叶银莲花组(Sect. Begoniifolia)等3分类群作为3亚组置于心皮无毛、花柱钩状弯曲、瘦果无肋的草玉梅组(Sect. Rivularidium)中。这与上述根据心皮和瘦果的形态特征划分组级分类群等的情况大不相同。   银莲花属
  Anemone L. Sp. Pl. 528. 1753; DC. Syst. 1: 188. 1817; et Prodr. 1: 16. 1824; Spach, Hist. Nat. Veg. Phan. 7: 242. 1839; Hook. f. & Thoms. Fl. Ind. 19. 1855; Prantl in Bot. Jahrb. 9: 224. 1887; Jancz. in Rev. Gén. 4: 241. 1892; Finet & Gagnep. in Bull. Soc. Bot. France 51: 56. 1904; Ulbr. in Bot. Jahrb. 37: 172. 1905; Juz. in Fl. URSS 7: 236. 1937; W. T. Wang in Fl. Reip. Pop. Sin. 28:1. 1980; Tamura in Acta Phytotax. Geobot. 12: 178. 1991; et in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17aIV: 324. 1995; Ziman et al. in J. Jap. Bot. 83(3): 127. 2008. Lectotype: A. coronaria L. (Britton & Brown, 1913; Javis et al., 1993)
  多年生草本植物。根状茎通常圆柱形,粗或细,具密集或疏离的节,稀呈块状。茎无叶,为花葶。叶均基生,少数或十数枚,稀不存在,为单叶,掌状分裂,稀不分裂,稀为三出复叶或羽状复叶(此时叶脉仍为掌状)。聚伞花序顶生,不分枝或1~4回分枝;总苞苞片3,稀2,4或更多,有柄或无柄。花萼片(4~) 5~6 (~15~20),花瓣状,花瓣不存在;雄蕊多数,通常无毛,花丝丝形或条形,花药内向或侧向;心皮少数至多数,子房被柔毛,有时无毛,花柱通常无毛,短或长,有时不存在,柱头不明显或明显,呈球形、卵球形或椭圆形、稀长圆形。瘦果无柄或有柄,不扁,呈卵球形等形状,有时两侧扁,甚至扁平,且在每侧具翅,通常无肋,稀具1条或4条肋;宿存花柱直或弯曲,或不明显。染色体基数x=8或7。
  本属有170余种,广布于世界各大洲,但多数种分布于亚洲,尤其是东亚。中国约有86种(包括59特有种),除在海南省没有分布外,广布于其他各省、区,多数种(约60种)分布于西南山区,在甘肃、陕西一带有10~15种,在新疆和内蒙古各有5~6种,在东北有13种,在湖南、湖北各有5~7种,向东种数逐渐减少,在华东沿海诸省和台湾地区,各有1~2(~3)种,在福建只有1变种(A. hupehensis var. japonica)。
  本文中,为了节省篇幅,只对刚恢复种级分类群的小花草玉梅(A. barbulata)、新提升到种级分类群的条叶银莲花(A. sino-linearis)和12新种写出形态描述,对其他71种(过去发表的种)则未写出形态描述。
  Anemone glaucifolia Franch. (Franchet, 1886)的叶羽状分裂,且具羽状脉,以及花的形状和颜色均与罂粟科植物十分相似。其花粉较大,具12条散沟和刺状纹饰,是高度特化特征(Xi & Chang, 1964)。在银莲花属的羽裂银莲花(A. subpinnata W. T. Wang), 以及毛茛科其他属如白头翁属的细叶白头翁(Pulsatilla turczaninoviiKryl. & Serg.) (Wang, 1980),铁线莲属的钝萼铁线莲(Clematis peterae Hand.-Mazz.) (Wang, 2003),毛茛属的宽翅毛茛(Ranunculus platyspermus Fisch.) (Wang, 1995)等诸种的叶均为羽状复叶,但叶脉却是掌状三出脉,不是羽状脉。银莲花属植物的花粉较小,多具3沟,只有3种[小花草玉梅(A. barbulata)、草玉梅(A. rivularis)和A. griffithii]的花粉具12條散沟,但无刺状纹饰(Xi & Chang, 1964),而与A. glaucifolia的花粉不同。由于A. glaucifolia具有上述诸特化形态特征,显示出与银莲花属存在重要区别,因此我将Franchet根据A. glaucifolia建立的单种组Sect. Anemoclema (Franchet, 1886)提升为属级群,即罂粟莲花属[Anemoclema (Franch.) W. T. Wang]。此属可能是在从毛茛科演化出罂粟科的漫长地质历史时期中出现的一个分类群。此单种属包含的植物即为罂粟莲花[Anemoclema glaucifolium (Franch.) W. T. Wang] (Wang, 1980)。如前所述,Hoot et al. (1994)认为A. glaucifolia和A. begoniifolia与分布于非洲的A. caffra群的植物在亲缘关系上极为相近。最近,我还看到一篇关于银莲花族(Trib. Anemoneae)的分子系统学文章( Liu et al., 2018),在此文的树系图中,Anemoclema glaucifolium不是与银莲花属(Anemone)植物在同一演化分枝上相聚集,而是与铁线莲属(Clematis)和锡兰莲属(Naravelia)2属植物在同一分枝上聚在一起;而银莲花属(Anemone)植物则与獐耳细辛属(Hepatica)植物和白头翁属(Pulsatilla)植物分别在另外2分枝上相聚集。根据上述,本文认为Anemoclema是隶属铁线莲亚族(Subtrid. Clematidinae)的一个成员。   中國银莲花属有20余种可供药用,如鹅掌草(Anemone flaccida)具有镇痛、抗炎、抗动脉粥状硬化和抗肿瘤作用;多被银莲花(A. raddeana)具有镇静、抗菌、抗肿瘤和抗氧化作用;西南银莲花(A. davidii)具有抗菌和抗肿瘤作用;这些植物的活性成分为皂苷类化合物(Xie et al., 2016)。银莲花属多数植物有美丽的花,可供观赏,像秋牡丹(A. hupehensis var. japonica)在中国长江流域及以南的省、区广泛栽培,供观赏,其他种类如打破碗花花(A. hupehensis)、大火草(A. tomentosa)、草玉梅(A. rivularis)、卵叶银莲花(A. begoniifolia)、多被银莲花(A. raddeana)、展毛银莲花(A. demissa)等均具美丽的花,都值得引种栽培。
  组1. 草玉梅组
  Sect.Rivularidium Jancz. in Rev. Gen. de Bot. 4: 251. 1892; Ulbr. in Bot. Jahrb. 37: 196. 1905; Tamura in Sci. Rep. Osaka Univ. 16(2): 26.1967; et in Hiepko. Nat. Pflanzenftam., Zwei. Aufl. 17aIV: 29. 1995; W. T. Wang in Fl. Reip. Pop. Sin. 28: 22. 1980; Ziman et al. in J. Jap. Bot. 53(3): 148. 2008; Q. E. Yang in Plant Syst. Evol. 301: 421. 2015, p. p. excl. Subsect. Vitifoliis, Subsect. Rupicolis et Subsect. Begoniifoliis. Lectotype: A. rivularis Buch.-Ham. ex DC.
  基生叶数枚,具长柄,通常3全裂或3深裂。萼片5~8 (~16)。雄蕊花丝通常丝形;花粉多具12条散沟(中国植物)。心皮和瘦果多数,无毛,稀子房被柔毛,具较长弯曲或钩状花柱。染色体基数通常x=8。
  约13种,分布于东亚、中美洲和南美洲。中国有3种,广布于西部和北部。
  1. 小花草玉梅 图版1: 1-4
  Anemone barbulata Turcz. in Bull. Soc. Imp. Nat. Mosc. 10(6-8): 149. 1837; Forbes & Hemsl. in J. Linn. Soc. Bot. 23: 10. 1886. —A. rivularis Buch.-Ham. ex DC. var. barbulata Turcz. ex Maxim. in Acta Hort. Petrop. 11: 21. 1890; Fedts. in Bull. Pekiing Soc. Nat. Hist. 2(3): 94. 1928; W. T. Wang in Acta Phytotax. Sin. 12(2): 109. 1974. — A. rivularis ssp. barbulata (Turcz.) Ulbr. in Bot. Jahrb. 37: 234. 1905. Holotype 河北: 1830-1840, P. Kirilow s. n. (LE, non vidi).
  Ranunculus mollendorffii Hance in J. Bot. 17: 7. 1879. Holotype 北京:百花山,alt. 5-6 000 ft., 1875-08-19, O. Moellendoff s. n. (non vidi)。
  Anemone rivularis var. flore-minore Maxim. Fl. Tangut. 6. 1889; Fl. Tsinling 1(2): 281, fig. 242. 1974; Fl. Pl. Herb.Chinae Bor. Or. 3: 158. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 24. 1980; S. Y. He, Fl. Beijing, rev. ed., 1: 243, fig. 306. 1984; J. W. Wang in Fl. Hebei 1: 463. 1986; D. Z. Ma & H. L. Liu, Fl. Ningxia 1: 191. 1986; Y. Z. Zhao in Fl. Intramongol., 2nd ed., 2: 465, pl. 188: 3-4. 1991; X. Y. Yu, B. L. Li & Y. M. Li in Fl. Shanxi. 1: 613, pl. 390.1992; J. G. Liu in Fl. Xinjiang. 2(1): 279. 1994; L. H. Zhou in Fl. Qinghai.1: 339. 1997; Wang. Ziman & Dutton in Fl. China 6: 316. 2001; X. D. Li, Y. Y. Zan & S. Huang in Fl. Shenongjia 285. 2017; J. Q. Fu in Fl. Loess.-plat. Sin. 1: 424, pl. 84: 1-4. 2000. — A. flore-minore (Maxim.) Y. Z. Zhao, Class. Distr. Vasc. Pl. Inn. Mongol. 162. 2012. Described from Kansu, Shaanxi and Hebei Provinces, no specimen cited.   A. rivularis auct. non Buch.-Ham. ex DC.: Franch. In Nouv. Arch. Mus. Hist. Nat. Paris, ser. 2, 5: 170. 1883; X. W. Kung, Fl. Lanzhou. 251, fig. 186. 1962.
  多年生草本,高(20~) 40~125 cm。基生葉5~10,具长柄;叶片纸质,五角形,2~8 × 4~12 cm,基部心形,3全裂,中央全裂片宽菱形,3半裂,边缘有小裂片和牙齿,侧全裂片斜扇形或斜倒卵形,不等2深裂,两面疏被伏毛;叶柄长5~16 cm,只近顶端有疏毛。花葶通常1条,无毛;聚伞花序或具1花,长3~10 cm,或具3~4条分枝,同时1~2回分枝,长达38 cm,具10余花;总苞苞片3,具短柄,宽菱形,长(1.5) 4~7 cm,3全裂,全裂片狭披针形,边缘具小齿,有时2~3浅裂; 花梗长1.5~14 cm。花:萼片5(~6), 白色,椭圆形或狭倒卵形,稀狭卵形,5.5~8(~9) × 2.5~4 mm,顶端圆形或钝,内面无毛,外面顶端和基部被短柔毛,有3条基生脉。雄蕊约20,无毛;花丝狭条形,1~3 × 0.18 mm;花药淡黄色,长圆形,1~1.2 × 0.5~0.6 mm。心皮12~32,无毛;子房狭长圆形,2 × 0.5 mm;花柱长约1.5 mm,钩状弯曲。瘦果扁,近长圆形,4 × 0.6 mm,无毛;宿存花柱长约2 mm,钩状弯曲。花期6月至9月。
  分布于中国重庆、湖北西部、四川、青海东部、新疆西部、甘肃、陕西、宁夏、陕西、河北西部和北部、内蒙古西南部。生于山地草坡、灌丛或林边,在四川和甘肃南部海拔1 800~3 600 m,在陕西、山西一带海拔1 200~3 000 m,在河北西部海拔500~1 600 m。
  在银莲花属,以及毛茛科的其他多数属中,花均具5枚萼片,花具多于5枚萼片乃是后来发生的进化现象。因此,花具5枚萼片,特产于中国的小花草玉梅(A. barbulata)在演化水平上要比草玉梅(A. rivularis)原始;后者的花通常具7~8枚萼片,可能是从小花草玉梅衍生出的一种植物,自中国西南部山区向西南方向分布达印度半岛和斯里兰卡岛。
  本种的根、根状茎和全草可供药用,有健胃消食、散瘀消结之效,用于肝炎、筋骨痛、牙痛等症(谢磊等,2016)。
  标本登录 重庆:巫溪,陈跃东2220;城口,戴天伦101091。湖北:神农架,神农架队31378。四川:石棉,谢朝俊42563;小金,张秀实,任有铣6033;金川,李馨75087;德格,崔友文4481;甘孜,崔友文4291;茂县,何铸,周子琳13196;汶川,方文培1467;阿坝,汤宗孝1391;红原,郎楷永,李良千,费勇2018。青海:湟中,何廷农952;民和,刘尚武2767;大通,张志和4338。新疆:特克斯,周太炎等65-658。甘肃:文县,白水江队534;舟曲,王作宾7458;武山,黄河队甘一队2078;榆中,黄河队3224;天祝,何业祺4504;山丹,青甘队3326;肃南,祁连山,何业祺3572. 陕西:凤县,王作宾96-22;太白山,杨金祥940;佛坪,郭本兆1458;宁陕,孔宪武3010;岚皋,巴山队1816。宁夏:同心,于兆英1632;海源,黄河队5577。山西:垣曲,包世英272;霍州,黄河队846;交城,夏纬瑛1076;兴县,王作宾3061;芦芽山,Harry Smith 8154;宁武,山西队356;五台山,关克俭,陈艺林1410;应县,段长虹304。河北:武安,关克俭5772;内丘,刘心源571;阜平,阜平队74;涞源,刘继孟3027;昜县,刘继孟255;小五台山,黄秀兰等5384;涿鹿,杨朝广1554;平泉,王忠涛等140487;围场,承德队202。北京:百花山,金德福514;东灵山,北京草药队555。内蒙古:兴和,赵一之77-11;凉城,内大第1组 51。
  2. 草玉梅 图版2:1-5
  Anemone rivularis Buch.-Ham.ex DC. Syst. 1: 211. 1817; et Prodr. 1: 21. 1824; Hook. f. & Thoms. Fl. Ind. 23. 1855; et in Hook. f. Fl. Brit. Ind. 1: 9. 1872; Franch. in Bull. Soc. Bot. France 33: 366. 1886; et Pl. Delav. 10. 1887; Maxim. in Acta Hort. Petrop. 11: 20. 1890; Finet & Gagnep. in Bull. Soc. Bot. France 51: 72. 1904; Ulbr. in Bot. Jahrb. 37: 234.1905; Hand.-Mazz. in Acta Hort. Gotob. 13: 176.1939, p. p. excl. syn. A. barbulata Turcz.; Iconogr. Corm. Sin. 1: 727, fig. 1453. 1972; Fl. Tsinling. 1(2): 72. 1974, p. p.; W. T. Wang in Fl. Reip. Pop. Sin. 28: 22. 1980; et in Fl. Xizang. 2: 78. 1985; Y. K. Li in Fl. Guizhou. 3:76, pl. 31: 7-9. 1990; W. T. Wang, Vasc. Pl. Hengduan Mount.1: 510. 1993; L. H. Zhou in Fl. Qinghai.1: 339. 1997; W. T. Wang in Fl. Yunnan. 11: 189. 2000; et in High. Pl. China 3: 492, fig. 785. 2000; Wang, Ziman & Dutton in Fl. China 6: 316. 2001; S. H. Fu, Fl. Hubei.1: 351.2001; Ziman et al. in J. Jap. Bot. 53(3): 149. 2008; L. Xie in Medic. Fl. China. 3: 317. 2016. — A. rivularis ssp. eurivularis Ulbr. in Bot. Jahrb. 37: 234. 1905. Holotype Nepal: Chitong, F. Buchanan s. n. (non vidi).   A. Leveillei Ulbr. in Bot. Jahrb. 36, Beibl. 80: 5, fig. 6-9. 1905.
  A. saniculifolia Lévl. in Repert. Sp. Nov. 7: 383. 1909.
  A. esquirolii Lévl. & Van. in I. c. 8: 88. 1910.
  A. longipes Tamura in Acta Phytotax. Geobot. 15: 192. 1954.
  A. rivularis var. pilosisepala W. T. Wang in Guihaia 33(5): 585. 2013. Holotype 西藏:米林,多雄拉山,alt. 4 200 m, 2012-07-30, 楊永,刘冰等281(PE)。
  2a. var. rivularis
  2ai. f. rivularis
  分布于中国西藏南部(海拔2 700~3 900 m)、云南(1 600~4 000 m)、广西西部(920~2 000 m)、贵州(850~1 300 m)、湖北西部(1 400 m)、重庆、四川(1 200~3 000 m)、青海东南部(2 600~3 700 m)、甘肃西南部,以及不丹、尼泊尔、印度、斯里兰卡。生于山地草坡、灌丛、河滩、溪边、林边或林下。
  本种的全草可供药用,有清热解毒、活血舒筋、止咳祛痰之效,用于咽喉肿痛、咳嗽痰多、癗疠结核、疟疾、牙痛、慢性肝炎等症;有毒(谢磊等,2016)。
  标本登录 西藏:普兰,青藏队8489;聂拉木,青藏队5737;亚东,张经炜2486;曲水,付国勋83;工布江达,覃海宁161;林芝,西藏中草药队3058;米林,李勃生,程树志5583;波密,应俊生,洪德元65-25;江达,青藏队11940。云南:德钦,俞德浚8347;中甸,俞德浚11815;维西,王启无64064;丽江,俞德浚15038;剑川,秦仁昌23065;兰坪,蔡希陶53783;洱源,汤宗孝291;大理,王启无63119;镇康,俞德浚17215;景东,周仕顺1452;昆明,钟观光2025;东川,蓝顺彬99;昭通,蔡希陶50870;巧家,孙必兴918;建水,蔡希陶57137;文山,蔡希陶51742。广西:那坡,中植联广西队1674。贵州:兴义,贵州队6970;安龙,张志松,张永田5120;贵阳,安明态5086;雷山,简卓坡等50792;遵义,A. N. Steward, 焦启源48。湖北:神农架,236-6队2513。重庆:南川:金佛山,李国凤62667;巫山,杨光辉59001;巫溪,杨光辉58848;城口,戴天伦101177。四川:攀枝花,杨永等209;普格,管中天8076;西昌,赵良能2188;盐源,姜恕5907;木里,俞德浚5790;稻城,四川植物队1414;得荣,青藏队1716;巴塘,郎楷永,李良千,费勇2315;雅江,应俊生3043;九龙,应俊生3908;雷波,俞德浚3863;屏山,俞德浚3366;汉源,刘振书498;大祥岭,Harry Smith 10205;石棉,谢朝俊41495;天全,胡文光,何铸10156;泸定,张永田,郎楷永27;康定,关克俭,王文采512;宝兴,俞德浚2294;小金,经济植物队59-195;丹八,D. E. Boufford等37950;甘孜,胡文光13019;德格,崔友文4494;金川,李馨77285;马尔康,李馨70594;黑水,经济植物队59-1394;若尔盖,四川植物队10246;汶川,汪发瓒21098;松潘,经济植物队59-1612;平武,鲁丽敏342;茂县,何铸,周子林12651。青海:囊谦,藏药队1184;玉树,?214;兴海,何廷农148;西宁,钟补求8142;门源,青藏队2547。甘肃:天祝,?556。
  2aii. 黄花草玉梅 (变型)
  f.lutea Ulbr. ex Hand.-Mazz. in Acta Hort. Gotob. 13: 176. 1939. Type 四川:松潘,alt.3 000~3 100 m, Harry Smith 2218 (holotype), 2659 (paratype)(non vidi)。
  本变型的萼片黄色,而与具白色萼片的模式变型不同。
  特产于中国四川松潘。生于海拔3 000~3 100 m山区。
  2b. 大理草玉梅 (变种)
  var. daliensis X. D. Dong & L. Yang in Bull. Bot. Res. Harbin 20(3): 248. 2000. Type云南:大理,alt. 2 900 m, 1998-05-20, 杨林980011 (holotype, non vidi), 980010 (paratype, non vidi.)。
  本变种的花具14~20枚淡红色萼片,而与模式变种不同。
  特产于中国云南大理。生于海拔2 900 m一带山地。
  3. 九龙银莲花 图版3
  Anemone jiulongensis W. T. Wang, sp. nov. Type 四川(Sichuan):九龙县,纳布厂,磨子沟 (Jiulong Xian, Nabuchang, Mezi Valley), alt. 3 817 m,山坡,花白色(on slope, fls. white),2018-07-20,张树仁,杨杰,张薇 (S. R. Zhang, J. Yang & W. Zhang) GGS 10336 (holotype and isotype)。   Perennial herbs 16-25 cm tall. Basal leaves5-6, long petiolate; blades papery., ovate 1.4-3.4 × 1.8-3.6 cm, at base shallowly cordate or subtruncate, 3-sect, central segenent rhombic,3-lobed, with lobes 2-3-lobulate, lateral segments smaller, flabellate-rhombic, unequally 3-lobulate, with lobules 2-3-dentate or 2-3-denticulate, adaxially appressed-puberulous, abaxially on prominent nerves with short hairs; petioles 3-9 cm long. Scapes 5-6, 9-15 cm long, sparsely puberulous; cymes terminal, 1-flowered; involuctral bracts 3, lanceolate, 1 cm long, entire, undivided, rarely 2-parted; pedicels (0.8-) 2.5-11 cm long, appressed-puberulous. Flower: Sepals 5, white, obovate or broad-oblanceolate, 8-10 ×4-6 mm, apex rounded. Stamens ca. 30, white, glabrous; filaments linear ca. 2.6 × 0.4 mm,1-nerved; anthers elliptic, ca. 1 × 0.4 mm; carpels ca. 3.6 mm long, ovaries ca. 2 mm long, densely villous, styles ca. 1.6 mm long, glabrous, apex recurved.
  特产于中国四川九龙。生于山坡上,海拔3 817 m。
  4. 细裂银莲花   图版2: 9-13
  Anemone filisecta C. Y. Wu & W. T. Wang in Acta Phytotax. Sin. 12(2): 165. 1974; W. T. Wang in Fl. Reip. Pop. Sin. 28: 24, pl. 9: 9-13. 1980; et in Fl. Yunnan. 11: 189. 2000; Wang, Ziman & Dutton in Fl. China 6: 316. 2001; Ziman et al in J. Jap. Bot. 83(3): 149. 2008. —Anemonidium filisectum (C. Y. Wu & W. T. Wang) Starod. in Vetren itsy Sist. Evo. 1. 119.
  1991. Holotype 云南:景洪,alt. 750 m,河岸, 1936-11,王啟无80898 (PE).
  特产于中国云南景洪。生于山地河岸,海拔750 m。
  组2. 西南银莲花组
  Sect. Anemonanthea DC. Syst. 1: 196.1817; et Prodr. 1: 18. 1824; Prantl in Bot. Jahrb. 9: 247. 1887; Ulbr. in Bot. Jahrb. 37: 191. 1905; W. T. Wang in Fl. Reip. Pop. Sin. 28: 8. 1980; Tamura in Hiepko, Nat. Pflanzenfam., Zwei Aufl. 17aIV: 338. 1995; Ziman et al. in J. Jap. Bot. 79(1): 46. 2004, et 83(3): 144. 2008. Lectotype: A. nemorosa L.
  Sect. Sylvia Gaudin, Fl. Helvet. 3: 490. 1828.
  Anemonoides sect. Umbrosa Starod. in Bot. Zhurn. 74: 1346. 1989. Type: A. umbrosa (C. A. Mey.) Holub.
  Sect.Rosulantes Ziman & Kadota in J. Jap. Bot. 79(1): 48 et (3): 203. 2004. Type: A. stolonifera Maxim.
  基生叶1~5,稀不存在,通常为单叶,3全裂,稀为三出复叶。聚伞花序有1~3 (~4)花;总苞苞片具柄,3全裂或为三出复叶。萼片5~15。雄蕊花丝丝形;花粉具3沟,稀具散沟(A. griffithii)。心皮无柄,花柱比子房稍短,柱头常明显。瘦果通常不扁。染色体基数x=8。
  约有32种,广布于北温带地区。中国有14种,多数分布于西南地区,数种分布于东北,在陕西一带只有2种。
  系1. 西南银莲花系
  Ser.Hylalectron (Irmisch) Ulbr. in Bot. Jahrb. 37: 193, 213. 1905; Tamura in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17aIV: 338. 1995. —Sect. Hylalectron Irmisch in Bot. Zeit. 14: 19. 1856. — Ser. Anemonanthea. Ziman et al. in J. Jap. Bot. 79(1): 47. 2004. Type: A. nemorosa L.   Anemonoides sect. Anemonoides subsect. Altaicae Starod. in Bot. Zhurn. 74: 1346. 1989. —Ser. Altaicae (Starod.) Ziman et al. in J. Jap. Bot. 79(1): 47, 65. 2004. Type: A. altaica (C. A. Mey.) Holub.
  Ser. Rosulantes. Ziman et al. in I. c. 48.
  Ser. Exiguae Ziman et al. in I. c. (1): 48 et (5): 281. Type: A. exigua Maxim.
  萼片5~15,水平开展,白色,有时蓝色或粉红色。雄蕊花丝丝形。
  约30种。同组分布。中国有13种。
  5. 西南银莲花 图版4:1-3
  Anemone davidii Franch. in Nouv. Arch. Mus. Hist. Nat. Paris, ser. 2, 8: 185. 1886; Pl. David. 2: 3. 1888; Ulbr. in Bot. Jahrb. 37: 233. 1905; Hand.-Mazz. in Acta. Hort. Gotob. 13: 175. 1939; Iconogr. Corm. Sin. 1: 724, fig. 1447. 1972; W. T. Wang in Acta Phytotax. Sin. 12(2): 109. 1974; et in Fl. Reip. Pop. Sin. 28: 8, pl. 1: 1-3. 1980; Y. K. Li in Fl. Guizhou. 3: 78, 1990; W. T. Wang in Fl. Yunnan. 11: 185. 2000; et in High. Pl. China 3: 488, fig. 777. 2000; K. M. Liu, Fl. Hunan 2: 658. 2000; Wang, Ziman & Dutton in Fl. China 6: 513. 2001; S. X. Fu, Fl. Hupeh. 1: 349. 2001; Ziman et al. in J. Jap. Bot. 79(3): 204. 2004, et 83(3): 146. 2008; L. Xie in Medic. Fl. Pl. 3: 304. 2016. — A. stolonifera Maxim. var. davidii (Franch.) Finet & Gagnep. in Bull. Soc. Bot. France 51: 72. 1904. —Anemonoides davidii (Franch.) Starrod, Ventrenysty 123. 1991. Holotype 四川:寶兴,1869-04, A. David s. n. (P, non vidi; photo, PE)。
  A. petiolulata Pei in Contr. Biol. Lab. Sci. Soc. China. Bot. ser. 9: 2, fig. 1. 1933. Holotype四川:南川,1930-05-27, H. F. Chang 273 (NAS).
  分布于中国西藏东南部(海拔2 300 m)、云南(1 400~3 500 m)、四川(1 000~3 000 m)、重庆、贵州(600~2 500 m)和湖北西部(650~2 000 m)。生于山坡草地、灌丛或林中。
  本种的根状茎可供药用,有活血止痛,祛瘀消肿之效,用于跌打损伤、风湿疼痛、腰肌劳损等症(谢磊等,2016)。
  标本登录 西藏:墨脱县,青藏队3976。云南:中甸,俞德浚11193;维西,王启无64619;怒江,澜沧江分水岭,G. Forrest 26368;大理,王汉臣4036;彝良,董洪进176;墨江,费德罗夫等147。四川:金阳,经济植物队59-3059;筠连,经济植物队59-129;马边,汪发瓒22973;峨边,俞德浚831;石棉,谢朝俊40308;峨眉山,杜大华234;荣经,傅德志84-285;天全,曲桂岭2437;都江堰,汪发瓒20843;安县,朱大海3731。重庆:南川,金佛山,李国凤60947;酉阳,西师生物系2461;武隆,昜思荣154;奉节,三峡队2841;城口,戴天伦105133。贵州:凯里,黔南队1221;江口,武陵山队1329;梵净山,黔北队1014。湖南:新宁,罗仲春1854;桑植,刘林翰910。湖北:兴山,蒋祖德,陶光复210;神农架,神农架队20816;竹溪,甘启良2775。
  6. 糙叶银莲花 图版4: 4
  Anmone scabriuscula W. W. Wang in Acta Phytotax. Sin. 12(2): 160. 1974; in Fl. Reip. Pop. Sin. 28: 10, pl. 1: 4. 1980; et in Fl. Yunnan 11: 187, pl. 54: 4. 2000; Wang, Ziman & Dutton in Fl. China 6: 314. 2001; Ziman et al. in J. Jap. Bot. 79(5): 283. 2004, et 83(3): 146. 2008. Holotype 云南:丽江,巨甸,新竹,alt. 2 900 m, 1958-09-11,王文采3123(PE)。
  特产于中国云南、丽江、巨甸。生于山地疏林下,海拔2 900 m。   7. 柏枝山银莲花 图版5: 1-3
  Anemone baizhishanica W. T. Wang, sp. nov. Holotype 重庆市:南川、柏枝山、三河坝(Chongqing: Nanchuan, Baizhi Shan, Sanheba),alt. 1 950 m,灌丛中,花白色(in bushes, fls. white),1986-05-09,金佛山考察队(Jinfo Shan Exped.)0147(PE)。
  Perennial herb ca. 16 cm tall. Basal leaves ca. 2, long petiolate; blades papery, cordate-pentagonal, ca. 2 × 4.5 cm, 3-sect, central segment broadly rhombic, shortly petiolulate (petiolule ca. 1.5 mm long), ca. 2 cm broad, 3-parted slightly exceeding the middle of segment, central lobe rhombic, 3-lobed, with lobules 3-5-denticulate, lateral lobes obliquely ovate, on upper margin few-denticulate, lateral segments sessile, obliquely flabellate, unequely 2-parted, with the upper lobes similar to the central segment, lower lobes much smaller; surfaces adaxially and abaxially sparsely appressed-puberulous; petioles ca. 6.5 cm long, only near apex puberulous.
  Scape singular, ca. 9.5 cm long, only near apex puberulous; monochasium ca. 7 cm long; involucral bracts 3, petiolate (petioles 1.7 cm long), with blades 3-sect, mostly similar to those of basal leaves in size and shape; pedicels 2-7 cm long, densely appressed-puberulous. Flower: Sepals 5, white, obovate, 18-19 × 11-12 mm,9-nerved, glabrous, at apex rounded. Stamens ca. 40, glabrous; filaments filiform, 3-3.5 mm long; anthers broad-oblong, 1 × 0.6-0.8 mm, apex obtuse. Gynoecium broad-conic, ca. 4 × 4.8 mm, consisting of ca. 70 glabrous carpels; ovaries narrow-ovoid, 0.6-0.8 × 0.3-0.5 mm; styles 0.3-0.5 mm long, adaxially with stigmatic tissue.
  特产于中国重庆市南川区柏枝山。生于山地灌丛中,海拔1 950 m。
  8. 匐枝银莲花 图版9: 4-5
  Anemone stolonifera Maxim. in Bull. Acad. Sci. St.-Petersb. 22: 225. 1876; Fl. Pl. Herb. Chinae Bor.-Or. 3: 154, pl. 65: 6-7. 1975; Liu & Hsieh in Fl. Taiwan 2: 480. 1976; W. T. Wang in Fl. Reip. Pop. Sin. 28: 10. 1980; Tamura & Shmizu in Satake et al. Wild Flow. Japan 2: 69. 1982; Yang & Huang in Fl. Taiwan, 2nd ed., 2: 511, pl. 237. 1996; Wang, Ziman & Dutton in Fl. China 6: 512. 2001; Ziman et al. in J. Jap. Bot. 79(3): 203. 2004, et 83(3): 146. 2008. — Anemonoides stolonifera (Maxim.) Holub in Folia Geobot. Phytotax. Praha 8: 156. 1973. Holotype 日本:Nambu, 1865, Tchonosky s. n. (LE, non vidi).
  A. siuzevi Kom. in Acta Hort. Petrop. 22: 262. 1903.
  分布于中国黑龙江、台湾,以及朝鲜、日本。生于山地阔叶林下。
  标本登录 黑龙江:黑河,?68;尚志,王光正s. n.(IFP);台湾:台中,杨宗愈等5646。
  9. 小银莲花 图版4: 5-8
  Anemone exigua Maxim. in Bull. Acad. Sci. St.-Petersb. 23: 306. 1877; et Fl. Tangut. 9, t.2: 7-15. 1889; Forbes & Hemsl. in J. Linn. Soc. Bot. 23: 11. 1886; Finet & Gagnep. in Bull. Soc. Bot. France 51: 65. 1904; Ulbr. in Bot. Jahrb. 37: 233. 1905; Hand.-Mazz. Symb. Sin. 7: 314. 1931; Iconogr. Corm. Sin. 1: 724, fig. 1448. 1972; Fl. Tsingling. 1(2): 278, fig. 239. 1974; W. T. Wang in Fl. Reip. Pop. Sin. 28: 11, pl. 1: 5-8. 1980; B. Z. Ding et al. Fl. Henan. 1: 478. 1981; D. Z. Ma & H. L. Liu, Fl. Ningxia 1: 191. 1986; X. Y. Yu, B. L. Li & Y. M. Li in Fl. Shanxi. 1: 611. 1992; L. H. Zhou in Fl. Qinghai. 1: 338, pl. 74: 5-8. 1997; W. T. Wang in Fl. Yunnan. 11: 187. 2000; et in High. Pl. China 3: 489, fig. 778. 2000; Wang, Ziman & Dutton in Fl. China 6: 313. 2001; Ziman et al. in J. Jap. Bot. 79(5): 281. 2004, p. p. excl syn. A. exigua var. shanxiensis B. L. Li & X. Y. Yu, et 83(3): 146. 2008. —Anemonoides exigua (Maxim.) Starod, Vetrenytsy 123. 1991. Type 甘肅:大通河,1873-05, N. M. Przewalski s. n. (holotype, LE; isotype, PE)。   A. takasagomontana Masamune in Not. Syst. Mus. Hist. Nat. Paris 6: 37. 1937. (fide Ziman et al., 2004)
  9a. var. exigua
  分布于中国云南西北部(丽江,海拔3 500 m)、四川西部(2 550~3 600 m)、青海东部、甘肃南部(1 600~3 500 m)、陕西南部(太白山,2 300~2 700 m)、山西中部(灵石,2 100 m)和台湾。
  Ziman et al. (2004)将Anemone takasagomontana Masamune归并于A. exigua Maxim.,于是形成本种在中国西部与台湾地区的一间断分布现象。在中国银莲花属中,水棉花(A. hupehensis f. alba)的分布区也有与小银莲花相似的间断分布的情况。
  标本登录 四川:康定,折多塘,T. Naito,郎楷永等523;金川,李谱雄10536;马尔康,吴中伦32528;理县,?872;茂县,何铸,周子林12663;九寨沟,韩裕丰等82-415;大邑,朱大海等20070689。青海:大通,刘继孟6534。甘肃:文县,白水江队445,5376;成县,王庆瑞11528;舟曲,王作宾14337;玛曲,白龙江队1602;迭部,白龙江队1834;卓尼,1885年,3—6月,G. N. Potanin s. n.; 洮河,1886年5—6月,G. N. Potanin s. n. 陕西:太白山,刘慎谔,钟补求2647。山西:灵石,唐进934。
  9b. 山西银莲花 (变种)
  var. shanxiensis B. L. Li & X. Y. Yu in Acta Phytotax. Sin. 27: 152. 1989, et in Fl. Shanxi. 1: 613, pl. 388. 1992; Wang, Ziman & Dutton in Fl. China 6: 313. 2001. Type 山西:灵石,太岳山,马和乡,红岩底村,alt. 2 100 m, 1987-05-28,余湘云87216(holotype, SXY),87215 (paratype, PE);同地,1987-05-28,李腊红87213(paratype, PE)。
  特产于中国山西灵石。生于山地桦树林下,海拔2 100 m。
  10.  乌德银莲花 图版6:13-15
  Anemone udensis Trautv. & C. A. Mey. Fl. Ochot. 6, t. 26. 1847; Maxim. Prim. Fl. Amur. 17. 1859; Finet & Gagnep. in Bull. Soc. Bot. France 51: 63. 1904; Ulbr. in Bot. Jahrb. 37: 221. 1905; Juz. in Fl. URSS 7: 250, pl. 15: 5a, b. 1937; Kitag. Lineam. Fl. Mansh. 214. 1939; Fl. Pl. Herb. Chinae Bor.-Or. 152. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 14, pl. 2: 13-15. 1980; S. H. Li, Fl. Liaonig 1: 511, pl. 210: 3-4. 1988; Wang, Ziman & Dutton in Fl. China 6: 314. 2001; Ziman et al. in J. Jap. Bot. 79(1): 59. 2004, et 83(3): 144. 2008; Y. Zhou, Colour. Ill. Pl. Wet Area. NE China 345. 2019. — Anemonoides udensis (Trautv. & C. A. Mey.) Holub in Folia Geobot. Phytotax. 8: 166. 1973. Holotype 俄罗斯(俄国):Khabarovski Reg., Udshoe, Siberi orient., 1844-06-18, Middendorf s. n. (LE, non vidi).
  分布于中国黑龙江东部以及俄罗斯远东地区。生于山地林边或灌丛中。
  标本登录 黑龙江:伊春,吴征镒134;牡丹江,J. Sato 7787。
  11. 阴地银莲花 图版6:10-12
  Anemone umbrosa C. A. Mey. in Ledeb. Fl. Alt. 2: 361. 1830; Kom. Fl. Mansh. 2: 263. 1903; Ulbr. in Bot. Jahrb. 37: 227. 1905; Kom & Alis. Key Pl. Far East. Reg. USSR 1: 541. 1931; Juz. in Fl. URSS 7: 248. 1937; Kitag. Lineam. Fl. Mansh. 214. 1939; Fl. Pl. Herb. Chinae Bor.-Or. 152. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 14, pl. 2: 10-12. 1980; S. H. Li, Fl. Liaoning 1: 511, pl. 210: 3-4. 1988; Wang, Ziman & Dutton in Fl. China 6: 315. 2001; Ziman et al. in J. Jap. Bot. 79(1): 62. 2004, et 83(3): 144. 2008; Y. Zhou, Colour Ill. Pl. Wet Area. NE China 343. 2019. —Anemonoides umbrosa (C. A. Mey.) Holud in Folia Geobot. Phytotax. 8: 166. 1973. Holotype 俄羅斯(俄国):“In sylvis umbrosissimis ad rivulum Grammatucha prope Riddersk Sibiriae altaicae”,Ledebour 795 (LE, non vidi).   分布于中国辽宁和吉林东部,以及朝鲜北部、俄罗斯西伯利亚地区。生于低山草地或林下,海拔200~600 m。
  標本登录 辽宁:草河口,赵大昌等601;丹东,鸡冠山,J. Sato 93;千山,刘慎谔等605。吉林:敦化,黄泥河,孔宪武1444,1507。
  12.  狭萼银莲花  图版7:1-4
  Anemone stenosepala W. T. Wang, sp. nov. Holotype 云南 (Yunnan):中甸,近仙人洞 (Zhongdian, near Xianrendong), alt. 3 400 m,沟边湿地,花白色 (in wet place by stream, fls. white),1998-08-24,杨亲二,孔宏智 (Q. E. Yang & H. Z. Kong) 98-110 (PE)。
  Very small perennial herbs, 2.5-10 cm tall. Rhizome oblique or horizontal, 6-9 mm long, ca. 2.5 mm across, putting forth a filiform stolon ca. 0.5 mm in diam. Basal leaves 2-3, long petiolate; blades cordate-pentagonal, 0.8-1.4 × 1.1-2 cm, 3-sect, central segment broadly rhombic, 0.6-1 cm broad, 3-lobed or 3-cleft, lobes 2-3-dentate, lateral segments slightly larger than central segment, also broadly rhombic, but oblique, unequally 2-cleft or 2-lobed; surfaces adaxially and abaxially appressed-pubescent; petioles 1-3.2 cm long, subglabrous. Scapes solitary, 1-7.5 cm long, above pubescent; cymes 1-flowered; involucral bracts 3, unequal in size and shape, petiolate, broadly triangular, rhombic or obliquely narrow-ovate, 4-5 × 2.5-5 mm,3-parted or unequally 2-lobed, lobes 1-2-dentate or entire, with petioles 2-3 mm long; pedicel 0.8-2 cm long, appressed-pubescent. Flower: Sepals 6, white, oblong, 4-4.5 × 1-1.5 mm,at apex slightly obtuse, 3-nerved, adaxially glabrous, abaxially with sparse short hairs. Stamens ca. 9, glabrous; filaments filiform, 0.3-1.5 mm long; anthers suborbicular, 0.5 × 0.7 mm, apex obtuse. Carpels 6-10, ca. 0.8 mm long, glabrous; ovaries long elliptic, 0.5 × 0.2 mm; styles 0.3 mm long. Achenes narrow-ovoid, 1-1.8 × 0.3-0.5 mm, densely pubescent; persistent styles straight or slightly recurved or nearly hooked, glabrous.
  特产于中国云南香格里拉县仙人洞。生于溪边湿地,海拔3 400 m。
  本种最小的植株只高2.5 cm。花极小,6枚白色长圆形萼片极小,长4~4.5 mm,宽1~1.5 mm,是中国银莲花属中最小的花。
  13. 黑水银莲花 图版6: 7-9
  Anemone amurensis(Korsh.) Kom. in Acta. Hort. Petrop. 22: 262. 1903; Kom. & Alis. Key Pl. Far East. Reg. USSR 1: 541. 1931; Juz. in Fl. URSS 7: 247. 1937; Kitag. Lineam. Fl. Mansh. 212. 1939; Fl. Pl. Herb. Chinae Bor.-Or. 150. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 15, pl. 2: 7-9. 1980; S. H. Li, Fl. Liaoning 1: 511, pl. 210: 1-2. 1988; W. T. Wang in High. Pl. China 3: 490, fig. 781. 2000; Wang, Ziman & Dutton in Fl. China 6: 314. 2001; Ziman et al. in J. Jap. Bot. 79(1): 55. 2004, et 83(3): 144. 2008; L. Xie in Medic. Fl. China 3: 310. 2016; Y. Zhou, Colour. Ill. Pl. Wet Area. NE China 340. 2019. — A. nemorosa L. ssp. amurensis Korsh. in Acta Hort. Petrop. 12: 292. 1892; Ulbr. in Bot. Jahrb. 37: 225. 1905. — Anemonoides amurensis (Korsh.) Holub in Folia Geobot. Phytotax. 8: 105. 1973.   分布于中国辽宁、吉林东部和黑龙江东部,以及朝鲜、俄罗斯远东地区。生于山地灌丛中或林下,海拔552~810 m。
  本种的全草可供药用,有发汗、强肝补肾之效,用于麻痹、胃痛、痛风、百日咳等症(谢磊等,2016)。
  标本登录 辽宁:桓仁,陈法东,胡清如87-05。吉林:敦化,傅德志83-09;长白山,方振富1331, 1351。黑龙江:小兴安岭,宋朝枢36。
  14. 阿尔泰银莲花 图版6: 4-6
  Anemone altaica Fisch. ex C. A. Mey. in Ledeb. Fl. Alt. 2: 362. 1830; Finet & Gagnep. in Bull. Soc. Bot. France 51: 64. 1904; Ulbr. in Bot. Jahrb. 37: 222. 1905; Juz. in Fl. URSS 7: 248. 1937; Fl. Tsiling. 1(2): 279, fig. 240. 1974; W. T. Wang in Fl. Reip. Pop. Sin. 28: 12, pl. 2: 4-6. 1980; B. Z. Ding et al. Fl. Henan. 1: 76. 1981; X. Y. Yu, B. L. Li & Y. M. Li in Fl. Shanxi. 1: 613, pl. 389. 1992; W. T. Wang in High. Pl. China 3: 489, fig. 779. 2000; Wang, Ziman & Dutton in Fl. China 6: 316. 2001; Ziman et al. J. Jap. Bot. 79(1): 65. 2004, et 83(3): 145. 2008; L. Xie in Medic. Fl. China 3: 306. 2016. — A. nemorosa L. ssp. altaica (Fisch. ex C. A. Mey.) Korsh. in Acta Hort. Petrop. 12: 29. 1892. — Anemonoides altaica (Fisch. ex C. A. Mey.) Holub in Folia Geobot. Phytotax. 8: 166. 1973. Holoetype 俄罗斯(俄国):“in montibus altaicis”, Schangin s. n. (LE, non vidi).
  分布于中国陕西南部、河南西部、山西南部和新疆,广布于亚洲北部和欧洲东部。生于山地灌丛、沟边或林下,海拔1 200~2 000 m。
  本种的根状茎可供药用,有化痰开窍,安神、祛风化湿等效,用于热痛神昏、癫痫、气闷耳聋、胸闷腹胀等症(谢磊等,2016)。
  标本登录 陕西:太白山,王作宾6399,傅坤俊2849;华阴,西安药检所生药室s. n.;华山,夏纬瑛,王振华105。河南:卢氏,朱长山2009;陕县,1959-05,?0629。山西:永济,刘天慰65;运城,黄河队423;垣曲,刘心源20066;阳城,包士英,严生俊171;翼城,黄河队23。新疆:尼勒克,1977-04-26,?517。
  15. 多被银莲花 图版6:1-3
  Anemone raddeana Regel in Bull. Soc. Nat. Mosc. 34: 16. 1861; Forbes & Hemsl in J. Linn. Soc. Bot. 23: 12. 1886; Finet & Gagnep. in Bull. Soc. Bot. France 51: 67. 1904; Ulbr. in Bot. Jahrb. 37: 220. 1905; Juz. in Fl. URSS 7: 249. 1937; Kitag, Lineam. Fl. Mansh. 213. 1939; Fl. Pl. Herb. Chinae Bor.-Or. 152. 1975; W. T. Wang in Fl. Reip. Pop. Sin 28: 12, pl. 2: 1-3. 1980; S. H. Li, Fl. Liaoning. 1: 509. 1988; Y. J. Zheng in Fl. Shandong 1: 35, fig. 27. 1997; W. T. Wang in High. Pl. China 3: 490, fig. 780. 2000; Wang, Ziman & Dutton in Fl. China 6: 315. 2001; Ziman et al. in J. Jap. Bot. 79(1): 68. 2004, p. p. excl. syn. A. raddeana var. lacerata Y. L. Xu, et 83 (3): 145. 2008; L. Xie in Medic. Fl. China 3: 308. 2016; Y. Zhou, Colour. Ill. Pl. Wet Area. NE China 336. 2019. — Anemonoides raddeana (Regel) Holub in Folia Geobot. Phytotax 8: 166. 1973. Holotype: Mandshuria ad fl. Amur montes Burejae, 1858-04, G. Radde s. n. (LE, non vidi).
  15a. var. raddeana
  分布于中國山东东部、辽宁、吉林和黑龙江,以及俄罗斯远东地区、朝鲜和日本。生于山地林边、林下,多石山坡或河滩上,海拔约800 m。   本种的根状茎可供药用,有祛风湿、消痈肿之效,用于风寒湿痹、四肢拘挛、风湿腰腿痛、关节疼痛、风寒或感冒等症(谢磊等,2016)。
  标本登录 山东: 昆嵛山,山东野生植物普查队0578。辽宁:宽甸,李冀云等1189;草河口,J. Sato 94。吉林:敦化,傅德志83-03;长白山,alt. 810 m,方振富1399;浩良河,丁托娅83-176。黑龙江:尚志,帽儿山,林大96级实习队96-02。
  15b. 龙王山银莲花 (变种)
  var. lacerata Y. L. Xu in Bull. Bot. Res. Harbin 13(2): 121. 1993; Wang, Ziman & Dutton in Fl. China 6: 315. 2001. Holotype 浙江、安吉、龙王山,alt. 950 m,阔叶林下,1991-04-04,徐耀良0879 (non vidi)。
  特产于中国浙江安吉龙王山。生于山地阔叶林下,海拔950 m。
  16.  三出银莲花 图版8:1-4;16: 9-11
  Anemone griffithii  Hook. f. & Thoms. Fl. Ind. 21. 1855; et in Hook. f. Fl. Brit. Ind. 1: 8. 1872; Bruhl. in Ann. Bot. Gard. Culc. 5: 75, t. 105A. 1896; Finet & Gagnep. in Bull. Soc. Bot. France 51: 71. 1904; W. T. Wang in Fl. Reip. Pop. Sin. 28: 11, pl. 11: 9-11. 1980; et in Fl. Xizang. 2: 78. 1985; Wang, Ziman & Dutton in Fl. China 6: 313. 2001; Ziman et al. in J. Jap. Bot. 79(5): 282. 2004, et 83(3): 146. 2008. — Anemonoides griffithii (Hook. f. & Thoms.) Holub in Folia Geobot. Phytotax. 8: 272. 1973. Holotype Bhutan: near Chuka, alt. 6000 ft. and Mishimi Hills, 1838, W. Griffith 1420 (K, non vidi).
  A. nanchuanensis W. T. Wang in Acta Phytotax. Sin. 12(2): 161, pl. 8: 1. 1974. Holotype 重慶:南川,小河乡,alt. 1 650 m, 1957-05-08,熊济华,周子林90728 (PE)。
  分布于中国西藏南部(海拔2 500~3 000 m)、四川中部(1 800~2 400 m)和重庆南部(1 800~1 950 m),以及不丹、尼泊尔和印度北部。生于山地林下、山坡草地或陡崖上。
  标本登录 西藏:聂拉木,樟木,张永田,郎楷永3412,3514;亚东,青藏补点队75-99。四川:都江堰,陈道纯30031;彭县,冯正波,朱大海2007-429;石棉,张树仁等2785。重庆:南川,金佛山,刘正宇5181,金佛山队149,182。
  17. 粗柱银莲花
  Anemone robustostylosa Miao in Acta Sci. Nat. Univ. Sunyats. 32(4): 56. 1993. Holotype 广西:乐业,雅长林场,alt. 950 m,路边,花紫色带红,1989-08-30,钟业聪890811 (SYS; photo, PE)。
  特产于中国广西乐业县雅长乡。生于低山路边,海拔950 m。
  系2.  反萼银莲花系
  Ser. Reflexae Ulbr. in Bot. Jahrb. 37: 194, 228. 1905; Tamura in Hiepko, Nat. Pflanzentam., Zwei. Aufl. 17aIV: 339. 1995; Ziman et al. in J. Jap. Bot. 79(1): 47. 2004. — Subsect. Reflexa (Ulbr.) Juz. in Fl. URSS 7: 250. 1937. —Anemonoides subsect. Reflexae (Ulbr.) Starod. Syst. & Evol. 123. 1991. Type: A. reflexa Steph.
  聚伞花序有1花。萼片5(~7),反折,绿白色,条形。雄蕊花丝狭条形。心皮子房密被柔毛。
  1种,分布于中国陕西(眉县、太白山、蓝田)、河南西部(嵩县)和吉林东部,以及朝鲜北部、俄罗斯远东和西伯利亚地区、蒙古和欧洲东部。
  18. 反萼银莲花 图版6:16-18
  Anemone reflexa  Steph. ex Willd. Sp. Pl. 2: 1282. 1797; Finet & Gagnep. in Bull. Soc. Bot. France 51: 61. 1904; Ulbr. in Bot. Jahrb. 37: 228. 1905; Juz. in Fl. URSS 7: 250, t. 15: 6. 1937; Fl. Tsinling. 1(2): 280. 1974; W. T. Wang in Acta Phytotax. Sin. 12(2): 109. 1974; Fl. Pl. Herb. Bor.-Or. 158. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 15, pl. 2: 16-18. 1980; in Acta Phytotax. Sin. 29(5): 162. 1991; et in High. Pl. China 3: 491, fig. 782. 2000; Wang, Ziman & Dutton in Fl. China 6: 315. 2001; Ziman et al. in J. Jap. Bot. 79(3): 197. 2004, et 83(3): 145, 2008; L. Xie in Medic. Fl. China 3: 312. 2016. — Anemonoides reflexa (Steph. ex Willd.) Holub in Folia Geobot. Phytotax. 8: 66. 1973. Lectotype 俄罗斯(俄国):“In Sibiriae regionibus Baicalensibus, Irkutsk”,Staphan & Laxmann s. n. (LE; Ziman et al., 2004)   同系分布。生于山谷灌丛中或河边湿地,在太白山分布于海拔1 200~1 400 m一带山地。
  标本登录 陕西:眉县,Zhu, Chen, Xu & Wang 2366; 太白山,王作宾6492,傅坤俊2849,杨金祥35。吉林:长白山,敖志文4108;集安,吉金祥633。
  组3.  钝裂银莲花组
  Sect. Himalayicae (Ulbr.) Juz. in Fl. URSS 7: 256. 1937; W. T. Wang in Fl. Reip. Pop. Sin. 28: 34. 1980; Tamura in Acta Phytotax. Geobot. 43: 179. 1991; et in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17aIV: 335. 1995; Ziman et al. in Edinb. J. Bot. 64(1): 51. 2007; et in J. Jap. Bot. 83(3): 138. 2008. — Sect. Pulsatilloides subsect. Brevistylae Ulbr. ser. Himalayicae Ulbr. in Bot. Jahrb. 37: 201. 1905. —Sect. Omalocarpus subsect. Himalayicae (Ulbr.) Tamura in Sci. Reip. Osaka Univ. 16(2): 27. 1967. — Pulsatilloides sect. Himalayica Starod. Syst. & Evol. 124. 1991. — Anamonastrum Holub sect. Anemonastrum subsect. Himalayicae (Ulbr.) Mosyaki in Photoneuron 55: 6. 2018. Lectotype: A. obtusiloba D. Don (Tamura, 1995)
  根状茎垂直。基生叶多为单叶,稀为三出复叶或羽状复叶;单叶叶片3全裂、深裂、浅裂或不分裂。聚伞花序多有1~2花;总苞苞片3,无柄。萼片5~8,白色,蓝色或黄色。雄蕊花丝条形;花粉具3沟。心皮子房密被柔毛,稀无毛,花柱稍短于子房,直,柱头不明显,稀明显。染色体基数x=7。
  约21种,分布于中国西部、北部和喜馬拉雅山区,以及阿富汗和吉尔吉斯坦。
  系1.  钝裂银莲花系
  Ser. Obtusilobae Ziman, Ehrendorfer & Bulakh in Edinb. J. Bot. 64(1): 56. 2007; et in J. Jap. Bot. 83(3): 147. 2008. Type: A. obtusiloba D. Don.
  基生叶具长柄,叶片心状五角形,3全裂,稀3深裂,稀卵形,不分裂,基部圆形。瘦果不扁或稍扁,无肋。
  约14种。
  19. 钝裂银莲花 图版9:1-3;12:11
  Anemone obtusiloba D. Don, Prodr. Fl. Nep. 194. 1825; Hook. f. & Thoms. Fl. Ind. 22. 1855; et in Hook. f. Fl. Brit. Ind. 1: 8. 1872; Franch. in Bull. Soc. Bot. France 33: 365. 1886; Finet & Gagnep in Bull. Soc. Bot. France 51: 62. 1904; Ulbr. in Bot. Jahrb. 37: 241. 1905; Lauener in Not. Bot. Gard. Edinb. 23: 181. 1960; W. T. Wang in Fl. Reip. Pop. Sin. 28:36. 1980; in Fl. Xizang. 2: 80, fig. 23:3. 1985; et in Fl. Yunnan 11: 193. 2000; Wang, Ziman & Dutlon in Fl. China 6: 326. 2001; Ziman et al. In Edinb. J. Bot. 64(1): 59, fig. 1. 2007; et in J. Jap. Bot. 83(3): 147. 2008; L. Xie in Medic. Fl. China 3: 328. 2016. — A.obtusiloba ssp. obtusiloba Bruhl in Ann. Bot. Gard. Culc. 5: 78. 1896. — A. obtusiloba ssp. genuina Ulbr. in Bot. Jahrb. 37: 242. 1905. — Anemonastrum obtusilobum (D. Don) Mosyokin in Photoneuron 55: 7. 2018. Lectotype Nepal: Gosainqsthan, Wallich s. n. (Ziman & Mosunkin, 2007). .
  A. obtusiloba var. chrysantha Ulbr. in Bot. Jahrb. 37: 242. 1906. Holotype 西藏:Kandang, Hans s. n. (non vidi)。
  A. obtusiloba var. villosa Marq. & Shaw in J. Linn. Soc. Bot. 48(321): 154. 1929. Holotype 西藏: Nyima La, alt. 3 960~4 270 m,杜鹃花灌丛,1924-06-22, F. Kingdon Ward 5821 (non vidi)。   19a. ssp. obtusiloba
  19ai. var. obtusiloba
  分布于中国西藏南部、云南西北部、四川西部、甘肃南部和新疆西部,以及尼泊尔、印度北部、巴基斯坦和中亚。生于高山草地或林下,海拔2 900~4 800 m。
  标本登录 西藏:吉隆,综考会队75-336;聂拉木,张永田,郎楷永3512;米林,西藏中草药队4029;昌都,青藏队1263。云南:中甸,1981-06-21,?67。四川:木里,郎楷永,李良千,费勇8;康定,折多山,郎楷永,李良千,费勇1063;二郎山,蒋兴麐s. n.;宝兴,C. S. Chang, H. Kim 等S10968;小金,覃海宁17013;道孚,林学系队01;金川,李馨77467;茂县,何铸,周子林30231;刷经寺,四川植物队9046。
  19aii. 光房银莲花 (变种) 图1
  var. leiovaria J. Q. Fu in Fl. Loess-plat. Sin. 1: 426, 611, pl. 84: 5. 2000. Type 青海:共和县,桠稍沟,alt. 3 560 m, 1963-06-30,张振万1913 (holotype, WUK, non vidi)。山西:五台山,龙泉寺,alt. 3 000 m, 1983-07-07,黄土高原植物考察队1413,1483 (paratypes, WUK, non vidi)。
  本变种的植株低矮,总苞苞片长圆形,不分裂,子房无毛,而与模式变种不同。
  分布于中国青海东部和山西五台山。生于海拔3 000~3 560 m山地一带。
  19b. 长柄钝裂银莲花 (亚种)
  ssp. dolichopoda W. T. Wang, ssp. nov. Type 西藏 (Xizang):吉隆縣,伦当(?)(Jilong Xian, Lundang ?) alt. 3 500 m, 山坡,花白色 (on slope, fls. white), 1975-07-16,青藏队草原组(Prairie Group, Qinghai-Xizang Exped.) 6953 (holotype, PE);吉隆县至邦兴(on the way from Jilong to Bangxing), alt. 2 800 m,花紫色(fls. purple), 1975-07-13,西藏队植被组(Vegetation Group, Xizang Exped.) 5233 (PE).
  A. ssp. obtusiloba differt plantis 36-60 cm altis, petiolis usque ad 29 cm longis, sepalis albis vel prupureis.Basal leaves ca. 2; blades cordate-pentagonal, up to 3.8 × 5 cm, 3-sect, adaxially glabrous, abaxially appressed-pubescent. Scapes 2-4, 26-50 cm long, sparsely pubescent; cymes 1-2-flowered; involucral bracts 3, sessile broad-rhombic, ca. 2 × 2 cm, 3-parted to near base; pedicels 4-10 cm long, with sparse hairs. Sepals white or purple, obovate, ca. 12 × 7 mm. Stamens 4-5 mm long, glabrous; filaments linear. Carpels 14-20; ovaries ellipsoid, 1.8-3 mm long, densely pubescent; styles 1-1.2 mm long, glabrous; stigmas inconspicuous.
  产于中国西藏吉隆。生于海拔2 800 m山地一带。
  19c. 光叶银莲花 (亚种) 图版12: 8
  ssp.leiophylla W. T. Wang in Fl. Reip. Pop. Sin. 28: 36, 350. 1980; Wang, Ziman & Dutton in Fl. China 6:326. 2001. — A. obtusiloba var. leiophylla (W. T. Wang) Ziman et al. in Edinb. J. Bot. 64(1): 64. 2007. Holotype 云南:贡山,alt.3 000 m,草甸,1937-07-12,俞德浚22076(PE)。
  分布于中国云南贡山和西藏墨脱。生于山地草坡、灌丛中、林边或沟边。海拔3 000~3 600 m。
  标本登录 云南:贡山,昆明植物所队9033,10025,青藏队8478,8484。西藏:墨脱,自那格至多雄拉,青藏队8362。
  19d. 镇康银莲花(亚种) 图版12: 7
  ssp.megaphylla W. T. Wang in Fl. Reip. Pop. Sin. 28: 36, 350. 1980; Wang, Ziman & Dutton in Fl. China 6: 326. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 65. 2007. Holotype 云南:镇康,雪山草坡,alt. 3 400 m,1938-07-24,俞德浚16956(PE)。   特产于中国云南镇康。生于山地草坡,海拔3 400 m。
  20. 泸定银莲花 图版10: 1-3
  Anemone ludingensis W. T. Wang, sp. nov. Holotype 四川 (Sichuan):泸定县,海螺沟,4号营地 (Luding Xian, Hailuogou, Camp no. 4), alt.3 738 m, 高山草甸 (on alpine meadow), 2018-07-15,杨永,李海文,杨智,王宇 (Y. Yang, H. W. Li, Z. Yang & Y. Wang) GGS 161 (PE)。
  Perennial herbs ca. 14 cm tall. Basal leaves ca. 5, long petiolate; blades papery, cordate-pentagonal, rarely suborbicular orbroad-ovate, 1.4-2.5 × 2.2-4.5 cm,3-sect, central segment broad-
  rhombic, 3-parted, lobes few-lobulate or few-denticulate, lateral segments obliquely flabellate, unequally 2-parted, with lobes lobulate and sparsely denticulate; surfaces both sparsely puberulous; petioles slender, 3.5-11 cm long, only near apex with sparse short hairs. Scapes slender, 5-12.5 cm long, subglabrous; cymes 1-flowered; involucral bracts 3, sessile, 1 larger, broad-rhombic, ca. 1.4 cm long and broad, 3-lobed, the second one similar to the first one, but smaller, ca. 1 cm long, the third one cuneate, ca. 1 cm long, unequally2-lobed; pedicels ca. 3.2 cm long, puberulous. Flower: Sepals 5, white, elliptic or obovate. 8-10 × 6 mm, glabrous, 5-6-nerved, apex rounded. Stamens ca. 40, 3-4 mm long, glabrous; filaments lanceolate-linear, 2.2-3.2 × 0.3-0.8 mm, 1-nerved; anthers yellowish, broad-oblong, 0.8-1 × 0.5-0.7 mm, apex obtuse. Carpels ca. 25; ovatries narrowly ovoid, 1-2 × 0.5-1 mm, densely pubescent; styles 1-1.4 mm long, glabrous; stigmas inconspicuous.
  產于中国四川泸定。生于高山草甸上,海拔3 738 m。
  21. 天全银莲花 图版11: 1-3
  Anemone patula Chang ex W. T. Wang in Acta Phytotax. Sin. 12(2): 169, pl. 9: 10. 1974; in Fl. Reip. Pop. Sin. 28: 40, pl. 10: 1-3. 1980; et in Vasc. Pl. Hengduan Mount. 1: 513. 1993; Wang, Ziman & Dutton in Fl. China 6: 526. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 65, fig. 2. 2007; et in J. Jap. Bot. 83(3): 147. 2008. —Anemonastrum patulum (W. T. Wang) Mosyakin in Phytoneruon 55: 7. 2018. Holotype 四川:天全,1936,曲桂龄2796 (PE)。
  21a. var. patula
  分布于中国四川西部。生于高山草地,灌丛中或冷杉林下,海拔3 600~4 350 m。
  标本登录 四川:金川,空卡山,第八森林调查队5022;松潘,大王山,alt. 4 000 m,成都生物所植被组30023;理县,大板昭,胡文光53-175,邓德之1215(均存SZ)。
  21b. 鸡足叶银莲花 (变种) 图版11: 4
  var.minor  W. T. Wang in Fl. Reip. Pop. Sin. 28: 41, 350, pl. 10: 4. 1980; et in Vasc. Pl. Hengduan Mount. 1: 513. 1993; Wang, Ziman & Dutton in Fl. China 6: 327. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 67. 2004. Type四川:宝兴,邓池沟,alt. 3 530 m,1933-06-26,杜大华4293 (holotype, PE);金川,卡萨,alt.3 950 m,高山灌丛,1958-07-31,第八森林调查队8835 (paratype, PE)。   分布于中国四川西部。生于山地灌丛或草地上,海拔3 500~4 000 m。
  标本登录 四川:马尔康,吴中伦32375;泸定,磨西,alt. 4 000 m,成都生物所植被组31254。
  22. 多果银莲花 图版14: 3-6
  Anemone polycarpa  W. E. Evans in Not. Bot. Gard. Edinb. 13: 154. 1921; Wang, Ziman & Dutton in Fl. China 6: 44. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 67, fig. 3. 2007; et in J. Jap. Bot. 83(3): 138. 2008. — A. rupestris Hook. f. & Thoms. ssp. polycarpa (W. E. Evans) W. T. Wang in Fl. Reip. Pop. Sin. 28: 43, pl. 11: 3-6. 1980; in Vasc. Pl. Hengduan Mount. 1: 513. 1993; et in Fl. Yunnan. 11: 198, pl. 58: 4-7. 2000. — Anemonastrum polycarpum(W. E. Evans) Mosyakin in Phytoneuron 55: 7. 2018. Holotype 云南:怒江,澜沧江分水岭,alt.12 000 ft., 1914-09-11, G. Forrest 13320 (E, non vidi)。
  分布于中国西藏东南部、云南西北部,以及不丹、尼泊尔、印度北部(fide Ziman et al., 2007)。生于高山草甸上,海拔3 600~4 800 m。
  标本登录 西藏:米林,西藏中草药队4029,李渤生,程树志5509,5530,5803;林芝,张经纬,王金亭190;波密,应俊生,洪德元65-846,生态室草原组14819。云南:贡山,俞德浚22236, 22374, 22585, 22678;俅江河谷,俞德浚19858;德钦,冯国楣5092, 5233, 6086, 6120, 6670。
  23. 波密银莲花 图版5: 4-6
  Anemone bomiensis W. T. Wang, sp. nov. Holotype 西藏 (Xizang):波密县,嘎隆拉雪山 (Bomi Xian, Galongla Snow Mountain), alt.3 734 m,高山草甸 (on alpine meadow), 2019-06-21, PE西藏考察队 (PE Xizang Exped.) 9195 (PE)。
  Small perennial herbs 4-6 cm tall. Basal leaves 3-4, long petiolate; blades chartaceous, cordate-pentagonal, 0.9-1.2 × 1.4-2 cm, 3-sect, central segment broadly rhombic, 3-cleft or 3-parted slightly exceeding the middle of segment, lobes once again 3-cleft, with lobules narrow-ovate, lateral segments similar to cetral segment, but slightly oblique and unequally 3-parted, with lobes 3-lobed; surfaces adaxially appressed-pubesoent, abaxially glabrous; petioles 1.6-2 cm long, subglabrous. Scapes 1-2, 3-5.5 cm long, subglabrous; cymes 1-flowered; involucral brancts 3, sessile, unequal, 6-10 mm long, 2 larger, broadly rhombic, 3-parted, 1 smaller, oblong, unequally 2-lobed or undivided; pedicels 0.8-1.5 cm long, above with sparse hairs. Flower: Sepals 5, white, obovate, rarely narrow-elliptic, 4-6 ×2-4 mm, glabrous, apex rounded. Stamens ca. 12, glabrous; filements white, linear, 1-2 mm long; anthers deep blue, broad-oblong, 0.8-1 mm long, apex obtuse. Carpels ca. 14; ovaries oblong-oblanceolate, 2 × 0.6 mm, on middle part pubescent; styles 1.2-1.3 mm long, glabrous, apex hooked; stigmas inconspicuous.
  產于中国西藏波密、嘎隆拉山北麓。生于高山草甸上,海拔3 734 m。
  24. 米林银莲花 图版1: 5-9
  Anemone milinensis W. T. Wang in Guihaia 33(5): 584, fig. 3: F-K. 2013, et 36, Suppl. 1: 80, fig. 3: F-K. 2016; et in High. Pl. China in Colour 3: 401. 2016. Holotype 西藏:米林,派区,多雄拉山,alt. 4 200 m,高山草地,花淡蓝紫色,2012-07-30,杨永,刘冰,林秦文,欧阳均,冯涛319 (PE)。   產于中国西藏米林和墨脱间的多雄拉山。生于高山草地或垫状灌丛草甸中,海拔3 900~4 200 m。
  标本登录 西藏:米林,派区,多雄拉山北坡,4 200 m,高山草甸中,1983-07-13,程树志,李渤生5517 (topotype);墨脱,那格,多雄拉山,alt.3 900 m, 1074-08-01, 青藏队3879。
  25. 岷山银莲花 图版12: 1-3
  Anemone rockii Ulbr. in Notizbl. Bot. Gart. Berl. 10: 876. 1929; Fl. Tsinling. 1(2): 284. 1974; W. T. Wang in Fl. Reip. Pop. Sin. 28: 40, pl. 8: 1-3. 1980; Wang, Ziman & Dutton in Fl. China 6: 327. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 69, fig. 10. 2007; et in J. Jap. Bot. 83(3): 148. 2008. — Anemonastrum rockii (Ulbr.) Mosyakin in Phytoneuron 55: 7. 2018. Type 甘肃:迭部,shimen之南,alt. 10 600 ft., 1925-06, J. F. Rock 12520 (lectotype, Ziman et al., 2007; isolectotype, PE);同地,Shimen, alt. 12 000 ft., 1925-07, J. F. Rock 13061 (syntype)。
  25a. var. rockii
  分布于中国西藏东部、云南西北、四川西北和甘肃南部以及尼泊尔 (fide Ziman et al., 2007)。
  25b. 多茎银莲花 (变种) 图版12: 9
  var. multicaulis W. T. Wang in Fl. Reip. Pop. Sin. 28:40, 350. 1980; et in Vasc. Pl. Hengduan Mount. 1: 513. 1993; Wang, Ziman & Dutton in Fl. China 6: 327. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 72. 2007. Type 四川:西昌,alt.2 900~3 100 m, 草地,1932-08-10,俞德浚1312 (holotype, PE), 1304 (paratype, PE)。
  特产于中国四川西昌。生于高山草地上,海拔2 900~3 100 m。
  25c. 巫溪银莲花 (变种) 图版12: 10
  var. pilocarpa W. T. Wang in Fl. Reip. Pop. Sin. 28: 40, 350. 1980; Wang, Ziman & Dutton in Fl. China 6: 327. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 71. 2007. Type 重庆:巫溪,alt. 2 300 m,草地,1935-08-27,曲桂龄2064 (holotype, PE);城口,alt. 2 100~2 500 m,1958-06,戴天伦100327,100797 (paratypes, PE)。
  产于中国重庆市北部。生于山地草坡,海拔2 100~2 500 m。
  26. 疏齿银莲花 图版13:1-3
  Anemone geum  Lévl. in Bull. Acad. Geogr. Bor. 25. 25. 1915; Hand.-Mazz. in Acta Hort. Gotob. 13: 177. 1939; Iconogr. Corm. Sin. 1: 729, fig. 1458. 1972; S. Y. He, Fl. Beijing, rev. ed., 1: 243, fig. 305. 1984; Wang, Ziman & Dutton in Fl. China 6: 327. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 72, fig. 6. 2007; et in J. Jap. Bot. 83(3): 148. 2008. — A. bonatiana Lévl. var. geum (Lévl.) Lévl. Cat. Pl. Yunnan 219. 1917. — Anemonastrum geum (Lévl.) Mosyakin in Phytoneuron 55: 7. 2018. Holotype 云南:Ta-hai-Tze, alt. 1 200 m, 1912-05, E. E. Maire s. n. (E; photo, PE)。
  A. obtusiloba D. Don ssp. ovalifolia Brühl in Ann. Bot. Gard. Culc. 5: 78., pl. 108B: 23, 27-30. 1896; Lauener in Not. Bot. Gard. Edinb. 23: 189. 1960; Fl. Tsinling. 1(2): 283, fig. 244. 1974; W. T. Wang in Fl. Reip. Pop. Sin. 28: 36, pl. 9: 1-3. 1980; et in Fl. Xizang. 2: 82. 1985; J. W. Wang in Fl. Hebei. 1: 465, fig. 468. 1986; D. Z. Ma & H. L. Liu, Fl. Ningxia. 1: 192. 1986; Y. Z. Zhao in Fl. Intramongol., 2nd ed., 2: 467, pl. 188: 1-2. 1991; X. Y. Yu, B. L. Li & Y. M. Li in Fl. Shanxi. 1: 620, pl. 393. 1992; J. G. Liu in Fl. Xinjiang. 2(1): 277, pl. 75:1-3. 1994; L. H. Zhou in Fl. Qinghhai 1: 340, pl. 74:9-12. 1997; W. T. Wean in Fl. Yunnan. 11: 194. 2000. — A. ovalifolia (Bruhl.) Hand.-Mazz. Symb. Sin. 7: 315. 1931. — A. geum Lévl, ssp. ovalifolia (Brühl.) Chaudhary in Bot. Zhurn. 73: 1188. 1988; L. Xie in Medic. Fl. China 3: 337. 2016. Based on var. geocharis Brühl and var. orthocaula Brühl.   A. obtusiloba var. geocharis Brühl in Ann. Bot. Gard. Culc. 5:78. 1896. Syntypes 甘肃:G. N. Potanin s. n. 四川:康定,A. E. Pratt176。西藏北部,N. M. Przewalski s. n. (non vidi.)。
  A. wardii Marq. & Shaw in J. Linn. Soc. Bot. 48: 154. 1929. Holotype 西藏:Doshong La. Pemako, 1924-10-25, F, Kingdon Ward 6262 (K. non vidi)。
  A. rupestris var. pilosa Marq. & Shaw in J. Linn. Soc. Bot. 48: 154. 1929. Holotype 西藏:Temo La, alt. 4 270 m, 1924-06-06, F. Kingdon Ward 5747 (K, non vidi)。
  A. obtusiloba ssp. ovalifolia Bruhl var. angustilimba W. T. Wang in Fl. Reip. Pop. Sin. 28: 37, 350, pl. 10: 5. 1980. Holotype 四川:雷波,黄毛梗,alt. 2 800 m, 1959-06-19, 四川经济植物队59-760 (PE)。
  分布于中国西藏南部和东部(海拔3 500~5 000 m)、云南北部(2 600~3 900 m)、四川西部(1 700~4 700 m)、青海、新疆中部和南部(4 200 m)、甘肃、宁夏(贺兰山,2 800 m)、陕西(陇县,1 900~2 300 m)、山西、河北北部(1 900~2 800 m),以及尼泊尔、印度北部(fide Ziman et al., 2007)。生于高山草地或灌丛中。
  本种的叶、花、果实及全草可供药用:叶、花和果实用于病后体温低、淋症、关节积液、黄水疮、咳嗽痰喘;全草用于止血(谢磊等,2016)。
  标本登录 西藏:普兰,青藏队8454;萨嘎,青藏队7107;仲巴,青藏队6536;吉隆,青藏队植被组5747;聂拉木,青藏队植被组4254,5647;定日,朱家柟21;亚东,青藏队2586;帕里,钟补求5774;拉萨,张永田,郎楷永1943;林周,傅国勋213;申扎,王金亭3607;班戈,陶德定10607;当雄,董洁华240;安多,陶德定10706;那曲,陶德定10538;索县,王金亭3210;嘉黎,陶德定10463;波密,应俊生,洪德元65-846;林芝,西藏中草药队3212;察隅,青藏队10580;芒康,青藏队11841;左贡,青藏队12137, D. E. Boufford, B. Bartholomew等40664, 40706;八宿,青藏队12231;察雅,Boufford & Bartholomew 等41233;类乌齐,Boufford等31995;昌都,综考会队76-55;贡觉,青藏队12556;江达,青藏队12462。云南:维西:王启无63858,64644,冯国楣4778;中甸,青藏队1259;洱源,金沙江队6323。四川:盐源,姜恕5997;得荣,D. E. Boufford等30944;木里,俞德浚5651,14589;乡城,D. E. Boufford等30425,30660;稻城,四川植物队1842;金阳,经济植物队59-3156;越西,俞德浚969;雷波,管中天8535;西昌,陈伟烈,陈家瑞5251;冕宁,西师生物系11932;Minya Konka雪山,J. F. Rock 17676;泸定,四川植物队6836;康定,方文培3553, Harry Smith 10462; 胡文光,何铸10482,11316;雅江,姜恕5140;理塘,姜恕5231;二郎山,经济植物队59-875;宝兴,俞德浚2376;张秀实,任有铣5494;金川,李馨77284,77816;道孚,张永田,郎楷永87;新龙,四川植物队6384;炉霍,D. E. Boufford等34708;甘孜,胡文光,何铸13020,13141;德格,四川植物队7256;色达,应俊生9076;壤塘,姜恕8945;阿坝,汤宗孝1407;马尔康,李馨70244,70596,70824,李谱雄10594;米亚罗,姜恕7153;理县,姜恕1164;茂县,何铸,周子林12710;巴朗山,汪发瓒21230;松潘,方文培4063,Harry Smith 2561, 2682;若尔盖,四川植物队10005,谭仲明93-34。青海:玉树,藏药队1215;德令哈,青甘队932;贵德,刘尚武3327;民和,何廷农781;共和,郭本兆,王遂义11921,12034;青海湖,关克俭77215;湟中,何廷农753;互助,郭本兆9328;海晏,钟补求8286;大通,张荣祖4467;门源,钟补求10086;新疆:乌鲁木齐,杨昌友214。甘肃:肃南,祁连山,何叶祺3656。山西:沁源,刘继孟1281。河北:小五臺山,王作宾595;东陵,段俊西s. n.。
  27. 嘎隆拉山银莲花 图版10: 4-6
  Anemone galonglashanica W. T. Wang, sp. nov. Holotype 西藏 (Xizang):墨脱县,嘎隆拉山之南 (Motuo Xian, in the south of Galongla Shan), alt. 4 150 m,灌丛草甸中,花白色 (in scrub-meadow, fls. white), 1982-08-14, 程树志,李渤生 (S. Z. Cheng & B. S. Li) 33 (PE)。   Perennial herbs 12-15 cm tall. Basal leaves4-10, 3-9.5 cm long, long petiolate, glabrous, rarely with lobes sparsely ciliolate; blades of larger leaves (with petioles 4-6.5 cm long) broadly ovate or cordate-pentagonal, 0.9-1.8 × 1.5-2.7 cm, at base cordate, 3-sect, segments shortly petiolulate or sessile, central segment broad-rhombic, 3-parted, with lobes 2-3-lobulate, 2-3-lobed or undivided, lateral segments obliquely flabellate, obliquely obovate or broad-rhombic, unequally 2-cleft or 2-3-lobed, with lobes 1-3-denticulate or entire, and blades of small leaves (with petioles 1.6-2.5 cm long) mostly broad-rhombic, 0.16-0.7×0.25-0.8 cm, 3-sect., with segments narrow-elliptic or linear, and 0.2-1.5 mm broad, entire. Scapes 4-7, 3.5-14 cm long, glabrous or near apex with a few hairs; cymes 1-3-flowered; involucral bracts 3, rarely 2, sessile, unequal, narrow-obovate, rhombic-obovate or obliquely oblong, 0.9-1.6×0.3-1 cm, 2-3-lobed or 2-3-cleft, rarely 3-parted, with entire narrow-ovate lobes, glabrous, rarely on some lobes sparsely ciliolate; pedicels 0.5-3 cm long, near apex with a few hairs. Flower: Sepals 5, white, narrow-obovate or obovate, 5-9×2.8-4.5 mm, glabrous, 5-nerved, apex truncate-rounded. Stamens ca. 40, glabrous; filaments linear, ca. 2.2×0.5-0.6 mm, 1-nerved; anthers broad-oblong, ca. 1 mm long, apex obtuse. Carpels 9-15; ovaries narrow-ovoid, ca. 2 mm long, abaxally densely pubescent; styles 0.2-1 mm long, glabrous, apex arcuate and abaxially with stigmas 0.2-0.5 mm long.
  产于中国西藏墨脱嘎隆拉山南麓。生于山地灌丛草甸中,海拔4 150 m。
  28. 短蕊银莲花 图版25: 5-7
  Anemone brachystema  W. T. Wang in Pl. Divers. Resour. 36(4): 450, fig. 1: E-G. 2014. Holotype 西藏:墨脱,多雄拉山之南,高山草甸,1980-06-24,生态室高原组10319 (PE)。
  特产于中国西藏墨脱多雄拉山南部高山草甸上。
  29.  玉龙山银莲花 图2
  Anemone yulongshanica W. T. Wang in Bull. Bot. Res. Harbin 16(2): 159, fig. 1: 1-4. 1996; et in Fl. Yunnan.11: 196. 2000; Wang, Ziman & Dutton in Fl. China 6: 325. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 84, fig. 14. 2007; et in J. Jap. Bot. 83(3): 148. 2008.  —Anemonastrum yulongshanicum (W. T. Wang) Mosyakin in Phytoneuron 55: 8. 2018. Holotype云南:麗江,玉龙雪山,母猪沟,溪边石上,1962-07-03,丽江植物园队10333 (KUN; photo, PE)。
  29a. var. yulongshanica
  特产于中国云南丽江玉龙雪山。生于山地溪边石上。
  29b. 截基银莲花 (变种) 图版11: 6-7
  var. truncata  (Comber) W. T. Wang in Bull. Bot. Res. Harbin 16(2): 159. 1996; Wang, Ziman & Dutton in Fl. China 6: 325. 2001; Ziman et al. in Edib. J. Bot. 64(1): 86, fig. 15. 2007. —A. coelestina Franch. var. truncata Comber in Not. Bot. Gard. Edinb. 18: 226. 1934. — A. obtusiloba ssp. ovalifolia var. truncata (Comber) W. T. Wang in Fl. Reip. Pop. Sin. 28: 37, pl. 10:6-7. 1980; in Vasc. Pl. Hengduan Mount. 1: 512. 1993; et in Fl. Yunnan 11: 196. 2000. Holotype 云南:丽江,玉龙雪山,alt.11 000 ft., 1910-05, G. Forrest 5640 (E, non vidi; photo, PE)。   产于中国云南丽江玉龙雪山和四川西南部(木里,美姑)。生于山地草坡或冷杉林下,海拔2 600~3 900 m。
  标本登录 云南:丽江,玉龙雪山,王启无71002,俞德浚15178,15595,赵致光30265.四川:木里,Ka-bu, alt. 3 900 m, 1937-05, 俞德浚5485;美姑,四川经济植物队59-1634。
  29c. 鹿马登银莲花 (变种)
  var. glabrescens  W. T. Wang in Acta Bot. Yunnan 30(5): 519, fig. 1: A-C. 2008. Type 云南:福貢,鹿马登乡,亚坪村,alt. 3 600~3 640 m, 2005-08, Gaoligong Shan Biocdiv. Survey 26397 (holotype, PE), 26453, 28024 (paratypes, CAS & PE);同地,利沙底,alt. 3 640 m, 2005-08-24, GSBS 28042 (paratypes, CAS & PE)。
  特产于中国云南福贡高黎贡山。生于海拔3 600~3 640 m一带山地。
  29d. 叶枝银莲花 (变种) 图版13: 10-13
  var. polysepala  (W. T. Wang) W. T. Wang, comb. nov. — A. obtusiloba ssp. ovalifolia var. polysepala W. T. Wang in Acta Phytotax. Sin. 12(2): 169. 1974; et in Fl. Reip. Pop. Sin. 28: 37, pl. 9: 10-13. 1980. Type 云南:维西,叶枝,alt. 3 780 m, 1940-05-29, 冯国梅4270 (holotype, PE), 4247 (paratype, PE)。
  特产于中国云南维西叶枝。生于海拔3 780 m一带山地。
  30. 微裂银莲花 图版13: 4-6
  Anemone subindivisa W. T. Wang in Acta Phytotax. Sin. 12(2): 173, pl. 10: 14. 1974; et in Fl. Reip. Pop. Sin. 28: 47, pl. 9: 4-6. 1980; in Vasc. Pl. Hengduan Mount. 1: 515. 1993; et in Fl. Yunnan. 11: 201, pl. 56: 7-8. 2000; Wang, Ziman & Dutton in Fl. China 6: 325. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 83, fig. 13. 2007; et in J. Jap. Bot. 83(3): 148. 2008. —Anemonastrum subindivisum (W. T. Wang) Mosyakin in Phytoneuron 55: 7. 2018. Type 四川:木里,米足嘎山,alt. 3 700~4 000 m, 1937-05-23, 俞德浚5756 (holotype, PE), 5635 (paratype, PE);同地,格布,alt. 3 400 m, 1937-05-19,俞德浚5499 (paratype, PE)。
  分布于中国云南西北部(洱源、丽江、中甸)和四川西南部(木里)。生于山坡草地以及冷杉林或云南松林下,海拔2 880~3 900 m。
  标本登录 云南:丽江,铁甲山,alt. 2 880 m,云南松林下,青藏队175;中甸,牧场上,冯国楣931。四川:木里,Shau-siang-liang-tze, alt.3 900 m, 1937-08,俞德浚7753;木里,米足嘎山,alt.3 050~4 800 m, 1928-06, J. F. Rock 16206, p. p。
  31. 三小叶银莲花 图版10: 7-9
  Anemone trifoliolata W. T. Wang, sp. nov. Holotype 四川:Djesi-La and Djesi-Longba, South of Tatsienlu, Fls. white, July 1929, J. F. Rock 17543 (PE)。
  Perennial herbs 7-14 cm tall. Basal leaves ca. 5, ternate, long petiolate; blades nearly deltoid, 2-3 × 2-4 cm; central leaflet shortly petiolulate, orbicular-ovate, 1.5-2.5 × 2-3 cm, at apex subrounded, at base subcordate or broadly cuneate, 3-cleft or 3-parted, with lobes incised and sparsely crenate-dentate, lateral leaflets slightly smaller, shortly petiolulate or sessile, unequally3-clert, all lobes and lobules imbricate among themselves, on both surfaces appressed-pubescent; petioles 2.5-4.5 cm long, densely spreading-villous. Scapes 2-6, 3-11 cm long, densely spreading-villous.; cymes 1-flowered; involucral bracts 3, sessile, unequal, narrow-obovate, narrow-lanceolate or ovate, 0.7-1.2 × 0.2-0.5 cm, entire, rarely on apex 3-denticulate; pedicels 2-7 cm long, densely appressed-pubescent. Flower: Sepals 5, white, of them 4 obovate and 1 broadly ovate, 9-12 × 6-8 mm, at apex rounded or obtuse, adaxially glabrous, abaxially sparsely appressed-pubescent. Stamens ca. 40, glabrous; filaments linear, 1-2 mm long. 1-nerved; anthers oblong, ca. 1 mm long, apex obtuse. Carpels ca. 9; ovaries narrow-oblong, 2 mm long, densely villous; styles 1.2 mm long, glabrous; stigmas inconspicuous.   产于中国四川康定之南的Djesi-La。
  32. 羽裂银莲花 图版13: 7-9
  Anemone subpinnata  W. T. Wang in Acta Phytotax. Sin. 12(2): 170, pl. 6: 7-9. 1974; in Fl. Reip. Pop. Sin. 28: 43, pl. 9: 7-9. 1980; et in Vasc. Pl. Hengduan Mount. 1: 514. 1993; Wang, Ziman & Dutton in Fl. China 6:326. 2001; Ziman et al. in Edinb. J. Bot. 69(1): 68, fig. 4. 2007; et in J. Jap. Bot. 83(3): 147. 2008. —Anemonastrum subp. innatum (W. T. Wang) Mosyakin in Phytoneuron 55: 8. 2018. Type 四川:木里,瓦欽,alt. 3 750 m,山坡草地,1937-06-21, 俞德浚6513 (holotype, PE), 6512 (paratype, PE)。
  产于中国四川西南部(木里、稻城)。生于山坡草地或灌丛中,海拔3 750~3 870 m。
  标本登录 四川:稻城,Bowa山,alt.3 870 m, 1998-07-04, D. E. Boufford, B. Bartholomew等28260。
  系2. 湿地银莲花系
  Ser. Rupestres Ziman, Ehrendorfer & Bulakh in Edinb. J. Bot. 64(1): 56. 2007.; et in J. Jap. Bot. 83(3): 147. 2008. Type: A. rupestris Hook. f. & Thoms.
  与Ser. Obtusilobae在亲缘关系方面极为相近,与后者的区别在于本系的瘦果两侧扁,并具肋。
  1种,分布见下。
  33. 湿地银莲花 图版14: 7-9
  Anemone neorupestris W. T. Wang, nom. nov. — A. rupestris Hook. f. & Thoms. Fl. Ind. 21. 1855, non A. rupestris Jacquem. ex Camb. 1844; et in Hook. f. Fl. Brit. Ind. 1: 9. 1872; Franch in Bull. Soc. Bot. France 33: 365. 1886; Forbes & Hemsl. in J. Linn. Soc. Bot. 23: 11. 1886; Ulbr. in Bot. Jahrb. 37: 242. 1905; Lauener in Not. Bot. Gard. Edinb. 23: 198. 1960; W. T. Wang in Fl. Reip. Pop. Sin. 28: 41, pl. 11: 7-9. 1980; in Vasc. Pl. Hengduan Mount. 1: 513. 1993; in Fl. Yunnan: 11: 198, pl. 58: 1-3. 2000; et in High. Pl. China 3: 496, fig. 793 2000; Wang, Ziman & Dutton in Fl. China 6: 323. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 86, fig. 16. 2007; et in J. Jap. Bot. 83(3): 148. 2008. — Anemonastrum rupestre (wall. ex Hook. f. & Thoms) Mosyakin in Phytoneuron 55: 7. 2018. Holotype: Sikkim, alt.4 570 m, J. D. Hooker s. n. (K, non vidi).
  33a. ssp. neorupestris
  33ai. var. neorupestris
  从傅德志教授的银莲花属(Anemone)属植物名录和Index Kewensis(Jackson, 1895),我得知于1844年已发表了一银莲花属植物,名为Anemone rupestris Jacquem. ex Camb. (in v. Jacquemont, Voyage dans l’Inde 4: 5. 1844)。因此,在1855年发表的Anemone rupestris Hook. f. & Thoms.是一晚出异物同名 (later homonym),根据《国际植物命名法规》(朱光华,2001)有关条文的规定是不合法的名称。本文将此晚出异物同名的种加词‘rupestris’改为‘neorupestris’。
  分布于中国西藏东南部(海拔3 700~4 200 m)、云南北部(2 700~4 600 m)和四川西部,以及不丹、尼泊尔和印度东北部。生于山地草坡、灌丛或溪边,海拔2 700~4 560 m。
  标本登录 西藏:米林,李渤生,程树志5604;波密,姚淦3273(NAS)。云南:德钦,杨亲二98-395;中甸,俞德浚12544,青藏队999,1259;丽江,
  王启无70522,71603,俞绍文65059;洱源,金沙江队6328;东川,蓝顺彬634。四川:松潘,汤宗孝634。   A. trullifolia Hook. f. & Thoms var. souliei Finet & Gagnep. in Bull. Soc. Bot. France 51: 62. 1904. Syntypes (均存P) 四川:康定,东俄洛,1891-07,Soulie 9, 17;同地,1892,Soulie 385;同地,1893-07-23,Soulie 688;巴塘与理塘间,1890-06-13,H. d’Orleans s. n. (non vidi)。
  39a. var. sino-linearis
  Perennial herbs. Rhizomes vertical, short and thick. Basal leaves 4-6, sessile, rarely petiolate; blades papery, broadly oblanceolate-linear or spatulate-oblanceolate, 3-12×0.7-2 cm, at apex3(-6)-dentate, rarely entire, adaxially appressed-puberulous, abaxially appressed-villous. Scapes 1-3(-7), 8-30 cm tall, puberulous; cymes1-flowered; involucral bracts 3, sessile, lanceolate, 0.7-1.5(-2.2) cm long, entire, rarely apically3-denticulate; pedicels sparsely puberulous. Flower: Sepals 5(-6), white, blue or yellow, obovate,10-13(-16)×5-7.5(-9) mm. Stamens numerous, 1.5-5 mm long, glabrous; filaments linear; anthers broadly oblong. Carpels 13-20; ovaries densely pubescent; styles short, glabrous; stigmas inconspicuous.
  从傅德志教授的银莲花属(Anemone)植物名录和Index Kewensis(Jackson, 1895),我得知于1841年已发表一银莲花属植物,学名为Anemone linearis Schlechtd. ex Pritz. (in Linnaea 15: 661. 1841.)。因此,如果将A. obtusiloba ssp. trullifolia var. linearis Bruhl提升为种级分类群,并根据此变种加词形成的种名必定是一不合法的晚出异物同名 (later homonym)。于是,本文将种名的种加词拟为‘sino-linearis’。
  分布于中国云南西北部、四川西部、甘肃南部、青海南部和西藏东部和南部以及不丹 (fide Ziman et al., 2007)。生于高山草地或灌丛中,海拔3 500~5 000 m。
  本种的全草、花和根可供药用:全草用于背部及脊椎骨痛;花和根用于慢性气管炎、末梢神经麻痹(谢磊等,2016)。
  标本登录 云南:中甸,俞德浚12092,冯国楣1556;德钦,姜恕9491。四川:稻城,D. E. Boufford, B. Bartholomew等28049,28161;乡城,D. E. Boufford, B. Bartholomew等28668;巴塘,D. E. Boufford, B. Bartholomew等36100;理塘,郎楷永,李良千,费勇2774;雅江,郎楷永,李良千,费勇2840;康定,郎楷永,李良千,费勇953,1014,关克俭,王文采603,1012;乾宁,四川植物队5391;泰宁,胡文光,何铸10886;道孚,姜恕2234;德格,四川植物队7156;色達,姜恕9077;金川,姜恕1118;马尔康,郎楷永,李良千,费勇2234;理县,姜恕1451;松潘,汤宗孝828;红原,姜恕7212;阿坝,汤宗孝1358。甘肃:玛曲,朱格麟4140;迭部,白龙江队1748;夏河,王作宾5726,7289;榆中,黄河队甘一队1712。青海:囊谦,杨永昌1100;玉树,1964-07,?302;同德,何廷农,B. Bartholomew等172。西藏:昌都,D. E. Boufford, B. Bartholomew等41128;芒康,青藏队11849;八宿,青藏队12241;林芝,吴征镒75-1162;米拉山,张永田,郎楷永1454;拉萨,张永田,郎楷永1941;羊八井,傅国勋173;南木林,西藏中草药队897;错那,吴征镒等75-913;帕里,钟补求5787;聂拉木,青藏队5738。
  39b. 钝齿条叶银莲花 (变种) 图版15:9-11
  var. holophylla  (Diels) W. T. Wang, comb. nov. — A. trullifolia Hook. f. & Thoms var. holophylla Diels in Not. Bot. Gard. Edinb. 5:263. 1912; Lauener in 1. c. 23: 197. 1960; W. T. Wang in Fl. Reip. Pop. Sin. 28: 47, pl. 12: 9-11. 1980; et in Fl. Yunnan. 11: 200, pl. 57: 10-12. 2000. — A. coelestina Franch var. polygyna Comb. f. holophylla (Diels) Comb. in Not. Bot. Gard. Edinb. 27: 226. 1924. — A. coelestina Franch. var. holophylla (Diels) Ziman & Dutton in Fl. China 6: 325. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 82, fig. 12. 2007. Holotype 云南:丽江,玉龙山,alt. 3 050~3 350 m, 1906-05, G. Forrest 2166 (E, non vidi)。   分布于中国云南丽江、四川木里和西藏东南部,以及尼泊尔和印度东北部 (fide Ziman et al., 2007)。生于山地草坡或沟边,海拔2 400~3 900 m。
  标本登录 云南:丽江,玉龙山,王启无71087,71602,俞德浚15208,青藏队304。四川:木里,俞德浚7248。
  组4. 银莲花组
  Sect. Omalocarpus DC. Syst. 1: 212. 1817; et Prodr. 1: 21. 1824; Hook. f. & Thoms. Fl. Ind. 23. 1855; Prantl in Bot. Jahrb. 9: 248. 1887;Ulbr. in Bot. Jahrb. 37: 209. 1905; W. T. Wang in Fl. Reip. Pop. Sin. 28: 47. 1980; Tamura in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17aIV: 336. 1995; Ziman et al. in J. Jap. Bot. 80 (5): 269. 2005, et 83(3): 137, 150. 2008. Lectotype: A. narcissiflora L.
  Anemonastrum Holub in Folia Geobot. Phytotax. 8: 158. 1973.
  Anemonastrum sect. Imbricata Starod. in Bot. Zhurn. 74: 1346. 1989. Type: A. imbricata (Maxim.) Holub.
  Sect. Leptothece W. T. Wang & Bing Liu in J. Syst. Evo. 46(5): 738. 2008. Type: A. xiaowutaishanica W. T. Wang & Bing Liu.
  根状茎垂直。基生叶3全裂或3深裂,稀为三出复叶。聚伞花序伞状,简单或复杂;总苞苞片无柄,3裂。萼片5或6,白色,蓝或紫色,稀黄色。雄蕊花丝条形。心皮花柱通常短,柱头不明显,稀明显。瘦果扁平,在每侧具翅;宿存花柱短。染色体基数x=7。
  约20种,分布于亚洲、欧洲和北美洲西部温带地区。中国有16种,广布于西部、北部和东北,多数种(11种)分布于西南山区。
  系1.  银莲花系
  Ser. Involucratae Ulbr. in Bot. Jahrb. 37: 209, 263. 1905-1906;Tamura in Sci. Rep. Osaka Univ. 16: 27. 1967; Ziman et al. in J. Jap. Bot. 80(5): 289. 2005. Type: A. narcissiflora L.
  伞状聚伞花序简单,有时只具1花。
  中国有12种。
  40a. 伏毛银莲花 (变种)
  Anemone narcissiflora L. var.  protracta Ulbr. in Bot. Jahrb. 37: 266. 1906; W. T. Wang in Fl. Reip. Pop. Pop. 28:47. 1980; J. G. Liu in Fl. Xinjiang 2(1): 279, pl. 74: 5, 1994; Ziman et al. in J. Jap. Bot. 80(5): 295. 2005. Described from Turkestan and no specimen cited.
  分布于中國新疆西部,以及哈萨克斯坦和塔吉克斯坦。生于山地草坡或云杉林中,海拔1 850~2 800 m。
  标本登录 新疆:昭苏,关克俭3472,李安仁,朱家柟10552,周太炎65-805。
  40b. 卵裂银莲花 (变种)
  Anemone narcissiflora L. var. sibirica (L.) Tamura in Acta Phytotax. Geobot. 17: 115. 1958; W. T. Wang in Fl. Reip. Pop. Sin. 28: 48. 1980; J. G. Liu in Fl. Xinjiang 2(1): 280, pl. 74: 4. 1994. — A. sibirica L. Sp. pl. 541. 1753; DC. Prodr. 1: 22. 1824; Y. Z. Zhao in Fl. Intramongol., 2nd ed., 2: 468, pl. 189: 4-5. 1991; et Class. Distr. Pl. Inn. Mongol. 163. 2012. Described from Altai.
  分布于中国新疆和内蒙古,以及蒙古和俄罗斯西伯利亚地区。生于山地草坡,海拔2 500~2 700 m。
  标本登录 新疆:乌鲁木齐,王治奎186;吐鲁番,李安仁,朱家柟5783.
  40c. 天山银莲花 (变种) 图版 16: 1-3
  Anemone narcissiflora L. var. turkestanica Schipcz. in Acta Hort. Bot. Univ. Jurjev. 13: 101. 1912; W. T. Wang in Fl. Reip. Pop. Sin. 28: 48, pl. 13: 1-3. 1980; J. G. Liu in Fl. Xinjiang 2(1): 280, pl. 74: 3. 1994. Described from Turkestan.   分布于中国新疆西部以及中亚地区。生于山地草坡,海拔1 900 m。
  标本登录 新疆:托里,关克俭2621,新疆师大队73152;乌鲁木齐,王多勤108,邹克荣140。
  40d. 长毛银莲花 (变种) 图版16: 8
  Anemone narcissiflora L. var. crinita (Juz.) Tamura in Acta Phytotax. Geobot. 17: 115. 1958; Fl. Pl. Herb. Chinae Bor.-Or. 3: 158. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 48, pl. 13: 8. 1980; S. Y. He, Fl. Beijing, rev. ed., 1: 245. 1984; J. G. Liu in Fl. Xinjiang 2(1): 279. 1994; Ziman et al. in J. Jap. Bot. 80(5): 295. 2005. — A. crinita Juz. in Fl. URSS 7: 274. 1937; J. W. Wang in Fl. Hebei. 1: 465. 1986; Y. Z. Zhao in Fl. Intramongol., 2nd ed., 2: 468, pl. 188: 6,8. 1991; et Class. Distr. Vasc. Pl. Inn. Mongol. 163. 2012. — A. narcissiflora L. ssp. crinita (Juz.) Kitag. Lineam. Fl. Mansh. 213. 1939. Lectotype: “Habitat in Sibira. Gmelin no. 71018 (Zimen et al., 2005)
  分布于中国河北北部、辽宁西部、内蒙古和新疆,以及蒙古、俄罗斯西伯利亚和远东地区。生于山坡草地、灌丛、林边或林下,海拔650~1 800 m。
  标本选录 北京:百花山,金德福151;灵山,杨朝广1542,1713。河北:赤城,大海坨山,陈岩,宋书银33;平泉,王忠涛等14-380;围场,李良千,王忠涛等49。内蒙古:克什克腾旗,杨雅玲等563;巴林右旗,翟秀丽,孙英宝2-019;巴林左旗,傅德志等81;乌兰浩特,傅德志等42;昭盟,蒙宁队473;牙克石,林学院队86-18;额尔古纳,王战1614;大兴安岭,J. Sato 86。
  40e. 贺兰山银莲花 (变种)
  var. alashanica Schipcz. in Acta Hort. Bot. Univ. Jurjev. 13(2): 100. 1912. — A. alashanica (Schipcz.) Borod.-Grabovsk. in Pl. Centr. Asia 12: 61, t. 5: 1-2. 2001; Y. Z. Zhao, Class. Distr. Vasc. Pl. Inn. Mongol. 163. 2012.
  Decribed from Helan Shan, Inn. Mongol.
  分布于中國内蒙古贺兰山和甘肃南部。生于山坡上或林下,海拔2 200~3 000 m。
  标本登录 内蒙古:贺兰山,何业祺2310,1860,阳文静,张彩飞90。甘肃:临潭,莲花山,黄河队洮河分队3411。
  41. 展毛银莲花 图版17: 4-6, 18: 1-2
  Anemone demissa  Hook. f. & Thoms. Fl. Ind. 23. 1855; et in Hook. f. Fl. Brit. Ind. 1: 9. 1872; Franch. in Bull. Soc. Bot. France 33: 367. 1886; Bruhl in Ann. Bot. Gard. Culc. 5: 81, t. 107. 1896; Ulbr. in Bot. Jahrb. 37: 267. 1906; Hand.-Mazz. in Acta Hort. Gotob. 13: 180. 1939; W. T. Wang in Fl. Reip. Pop. Sin. 28: 48, pl. 14: 4-6. 1980; in Fl. Xizang. 2: 82, fig. 22: 1. 1985; Y. Z. Zhao in Fl. Intramongol. 2nd ed., 2: 468, pl. 189: 1-3. 1991; L. H. Zhou in Fl. Qinghai. 1: 341. 1997; W. T. Wang in Fl. Yunnan. 11: 201. 2000; et in High. Pl. China 3: 497, fig. 796. 2000; Wang, Ziman & Dutton in Fl. China 6: 321. 2001; Ziman et al in J. Jap.Bot. 81(1): 1. 2006, et 83(3): 151. 2008. —A. narcissifolia L. var. demissa (Hook. f. & Thoms.) Finet & Gagnep. in Bull. Soc. Bot. Bot. France 51: 74. 1904. — Anemonastrum demissum (Hook. f. & Thoms.) Holub in Folia Geobot. Phytotax. 8: 105. 1973. Holotype: Sikkim: Lama Gingra, Sarnclery, alt. 16000 ft., 1849-07-15, J. D. Hooker s. n. (K, non vidi).   A. demissa var. yunnanensis Franch in Bull. Soc. Bot. France 33: 367. 1886; Pl. Delav. 10. 1889; W. T. Wang in Fl. Reip. Pop. Sin. 28: 51. 1980; et in Fl. Yunnan 11: 201, pl. 59: 1-2. 2000; Wang, Ziman & Dutton in Fl. China 6: 322. 2001; Ziman et al. in J. Jap. Bot. 81(1): 4. 2006. Syntypes 云南:大理,苍山,alt. 3 000~3 500 m, 1884-06, J. M. Delavay 154 (P; photo, PE), 989 (P, non vidi)。
  A. demissa var. grandiflora Marq. & Shaw in J. Linn. Soc. Bot. 48(321): 154. 1929.
  41a. var. demissa
  分布于中国云南西北部、四川西北部、甘肃西南部、青海南部、西藏东部和南部,以及不丹、尼泊尔和印度北部。生于山地草坡或疏林中,海拔3 200~4 600 m。
  本种的根状茎、叶和果实可供药用:根状茎用于疟疾、恶疮;叶和果实用于祛风除湿、防腐杀菌(谢磊等,2016)。
  标本登录 云南:德钦,王启无69430,70112;中甸,汤宗孝511;丽江,汤宗孝449。四川:木里,韩裕丰83-109;乡城,四川植被队953,3260;稻城,四川植被队1709;九龙,应俊生3806;雷波,管中天8532;泸定,朱大海3897;康定,Harry Smith 10610,关克俭,王文采1102;理塘,郎楷永,李良千,费勇2803;巴塘,D. E. Boufford, B. Bartholomew等36097;德格,四川植被队7129;甘孜,姜恕2418;壤塘,应俊生9010;宝兴,俞德浚2318;小金,张秀实,任有铣6197;金川,李馨77765;马尔康,李馨71611;茂县,汤宗孝91;黑水,姜恕1454;阿坝,汤宗孝56;红原,四川植物队9089;若尔盖,谭仲明93-36。青海:玉树,?374。西藏:江达,青藏队76-12491;察隅,青藏队73-467;贡觉,D. E. Boufford, B. Bartholomew等41076;芒康,青藏队11850;八宿,李渤生等7360;昌都,青藏队3608;类乌齐,杨竞生91-624;墨竹工卡,倪志诚6;米林,西藏中草药队3795;林芝,西藏中草药队3204;拉薩,张永田,郎楷永2024;朗县,西藏中草药队4466;聂拉木,张永田,郎楷永3978;吉隆,西藏中草药队664。
  41b. 矮展毛银莲花 (新变种) 图版8: 5-7
  var. humilis W. T. Wang, var. nov. Holotype
  四川 (Sichuan):九龙县,汤古乡,扎拖沟 (Jiulong Xian, Tanggu Xiang, Katuo Valley), alt.3 854 m,沼泽中,花白色 (in swamp, fls. white),2018-09-10,陈文莉,蒋善文,王继旋 (W. L. Chen, S. W. Jiang & J. X. Wang) 1588 (holotype, PE)。
  A var.demissa differt plantis humilibus ca. 11 cm altis, foliorum petiolis multo brevioribus 1-2.6 cm longis cum scapis appresso-villosis, et sepalis albis. In var. demissa, plantae vulgo 20-45 cm altae, foliorum petioli 9-15 cm longi cum scapis patule villosi, et sepala caerulea vel purpurea, raro alba sunt.
  此变种的植株矮,高约11 cm,叶柄短得多,长1~2.6 cm,与花葶均被贴伏长柔毛,萼片白色,而与模式变种不同。在模式变种,植株通常高20~45 cm,叶柄长9~15 cm,与花葶均被开展长柔毛,萼片呈蓝色或紫色,稀呈白色。
  特产于中国四川九龙。生于山地沼泽地,海拔3 845 m。
  41c. 宽叶展毛银莲花 (变种) 图版16:4-7
  var. major W. T. Wang in Fl. Reip. Pop. Sin. 28: 51, 351, pl. 13: 4-7. 1980; in Fl. Xizang 2: 83. 1985; in Vasc. Pl. Hengduan Mount. 1: 515. 1993; et in Fl. Yunnan 11: 203, pl. 59: 3. 2000; Wang, Ziman & Dutton in Fl. China 6: 322. 2001; Ziman et al. in J. Jap. Bot. 81(1): 5. 2006. Holotype 云南:维西,东竹岭,alt. 4 000 m,山坡,花淡蓝,1937-07-11,俞德浚8908 (PE). Paratypes(均存PE) 云南:维西,王启无63823,64472;德钦,俞德浚8735, 8745, 9842;中甸,俞德浚11745。四川:木里,俞德浚5657, 7076;冕宁,武素功2229;雷波,四川经济植物队843;鹧鸪山,李馨71687;绰斯甲,李谱雄10504。   分布于中国云南西部至东北部、西藏东部、四川西部和北部、甘肃西南部。生于山地草坡或疏林中,海拔2 300~4 700 m。
  标本登录 云南:贡山,青藏队8637;丽江,毛品一139,杨亲二,孔宏智98-477;宁蒗,汤宗孝1061;兰坪,1960-07,?9807;镇康,alt. 3 400 m,俞德浚117198;巧家,alt. 3 000 m,孙必兴1020。西藏:墨脱,alt. 3 200 m,综考会队76-332;米林,alt. 4 000 m,倪志诚等3023;错那,资源考察队1697。四川:美姑,经济植物队59-1622;九龙,应俊生3812;乡城,青藏队4089,D. E. Boufford, B. Bartholomew等28630;理塘,郎楷永,李良千,费勇2803;金川,田效文1010;黑水,经济植物队59-1345;松潘,1928-08,方文培4041;南江,光雾山,alt. 2 300 m,巴山队5685。甘肃:文县,alt. 3 300 m,白水江队5111。
  41d. 密毛银莲花 (变种)
  var.villosissima Bruhl in Ann. Bot. Gard. Culc. 5: 81. 1896; Hand.-Mazz. in Acta. Hort. Gotob. 13:180. 1939; W. T. Wang in Fl. Reip. Pop. Sin. 28: 51. 1980; in Fl. Xizang 2: 83. 1985, et in Fl. Yunnan 11: 203. 2000; Wang, Ziman & Dutton in Fl. China 6: 322. 2001; Ziman et al. in J. Jap. Bot. 81(1): 4. 2006. — A. demissa ssp. villosissima (Bruhl) Chaudkhary in Bot. Zhurn. 73(8): 196. 1988. HolotypeSikkim: Jogri, alt.13 000 ft., 1892-06-22, Gammie 195 (non vidi).
  A. demissa var. villosa Ulbr. in Bot. Jahrb. 37: 209. 1906.
  分布于中国云南西北部、西藏南部、四川西部、甘肃南部,以及尼泊尔、印度北部 (fide Ziman et al., 2006)。生于山地草坡、灌丛中或林中,海拔2 500~4 450 m。
  标本登录 云南:泸水,赵祖义120;福贡,Gaoligong Shan Biodiv, Survey 26430;德钦,王启无64972;中甸,冯国楣1248;丽江,俞德浚15055,王启无71592;大理,王启无63189。四川:木里,俞德浚5657,14759;雷波,四川经济植物队59-9251;马边,汪发瓒22034;九龙,俞德浚6829;宝兴,张秀实,任有铣5532;金川,李馨77550;茂县,何铸,周子琳12721;都江堰,汪发瓒20900。甘肃:舟曲,王作宾14828;临潭,黄河队3411。
  42. 粗壮银莲花
  Anemone robusta W. T. Wang in Acta Phytotax. Sin. 12(2): 173, pl. 10:15. 1974; in Fl. Reip. Pop. Sin. 28:53. 1980; in Vasc. Pl.Hengduan Mount. 1: 516. 1993; et in Fl. Yunnan. 11: 203. 2000; Wang, Ziman & Dutton in Fl. China 6: 320. 2001; Ziman et al. in J. Jap. Bot. 81(1): 5. 2006, et 83(3):151. 2008. Type 云南:维西,alt. 2 500~3 000 m,1935年6—9月,王启无67852 (holotype, PE), 63778 (paratype, PE)。
  分布于中国云南西北部(维西、中甸、德钦)和四川西南部(木里)。生于高山草甸上或灌丛中,海拔2 500~4 600 m。
  标本登录 云南:中甸,大雪山丫口,alt.4 300 m,杨亲二,孔宏智98-273;德钦,换夫坪,alt.3 600 m,王启无69018。四川:木里,三区,alt. 4 600 m, 青藏队14473。
  43. 太白银莲花 图版18: 3-4
  Anemone taipaiensis W. T. Wang in Acta Phytotax. Sin. 12(2): 174, pl. 10: 16. 1974; Fl. Tsinling. 1(2): 284. 1974; W. T. Wang in Fl. Reip. Pop. Sin. 28: 53, pl. 15: 3-4. 1980; Wang, Ziman & Dutton in Fl. China 6: 322. 2001; Ziman et al. in J. Jap. Bot. 81(1): 5. 2006; et 83(3): 151. 2008. Holotype 陜西:太白山,放羊寺,alt.3 200~3 600 m,1937-09,刘慎谔,钟补求958 (PE)。
  产于中国陕西南部秦岭山区(太白山,宝鸡)。生于高山草地、多石山坡或陡崖上,海拔2 900~3 700 m。
  标本登录 陕西:太白山,alt. 2 900~3 700 m,王作宾1777,刘慎谔,钟补求839,汪发瓒182,秦岭队10383,杨金祥910,Zhu-Chen-Xu-Wang 1380,2474;宝鸡,陕西中医研究所1978。   44. 银莲花 图版18: 5-9
  Anemone cathayensis Kitag. Lineam. Fl. Mansh. 213. 1939; Tamura in Acta Phytotax. Geobot. 17: 114. 1958; Iconogr. Gorm. Sin. 1: 730, fig. 1460. 1972; W. T. Wang in Fl. Reip. Pop. Sin. 28: 54, pl. 15: 5-9. 1980; S. Y. He, Fl. Beijing, rev. ed., 1:241, fig. 306. 1984; J. W. Wang in Fl. Hebei. 1: 466, fig. 469. 1986; X. Y. Yu, B. L. Li & Y. M. Li in Fl. Shanxi. 1: 620, pl. 394. 1992; W. T. Wang in High. Pl. China 3: 498, fig. 797. 2000; Wang, Ziman & Dutton in Fl. China 6: 321. 2001; Ziman et al. in J. Jap. Bot. 81(1): 7. 2006, p. p. excl. syn. A. cathayensis var. hispida Tamura et 83(3): 151. 2008; J. C. Zhao et al. Fl. Xiaowutaishan 1: 315, fig. 286. 2011; L. Xie in Medic. Fl. China 3: 334. 2016. Holotype 河北:小五臺山,1914, Y. Nagai 48(TI, non vidi)。
  A. demissa Hook. f. & Thoms. var. glabrescens Ulbr. in Bot. Jahrb. 37: 209. 1906; Hand.-Mazz. in Acta Hort. Gotob. 13: 181. 1939.
  A. narcissiflora L. var. pekinensis Schipcz. in Acta Hort. Bot. Univ. Jurejev. 13: 101. 1912.
  A. narcissiflora L. var. chinensis Kitag. in Rep. Ist Sci. Exp. Manch. 4(4): 17. 1935; Fl. Pl. Herb. Chinae Bor.-Or. 3: 158. 1975. —A. narcissiflora L. ssp. chinensis (Kitag.) Kitag. Lineam. Fl. Mansh. 213. 1939.
  44a. var. cathayensis
  分布于中国河北、山西和河南北部。生于山区草地、灌丛或林下,海拔1 000~2 500 m。
  本种的根可供药用,用于止血除湿、清热解毒(谢磊等,2016)。
  标本登录 河北:武安,alt. 1 400 m,关克俭76;内丘,刘瑛12965;涞源,河北农大队12357;涞水,alt. 1 000 m,刘继孟2190;小五台山,alt.2 000~2 390 m,王启无61344,杨朝广406,段俊喜5290;杨家坪,刘瑛10376;东陵,唐进2208;青龙,王忠涛等14-296;雾灵山,刘慎谔等4775,刘继孟59。北京:百花山,刘冰等686;小龙门,关克俭74-142;怀柔,海坨山,alt. 1 500 m,怀柔队391;东灵山,alt. 2 020 m,段俊喜92。山西:凌川,包士英,严生俊1332;沁源,刘继孟1416;霍县,黄河队710;介休,Harry Smith 5891,刘继孟2025。河南:林县,河南植物普查队59-4068。
  44b. 毛蕊银莲花 (变种) 图版18: 10
  var.hispida Tamura in Acta Phytotax. Geobot. 17: 114. 1958; W. T. Wang in Fl. Reip. Pop. Sin. 28: 54. 1980; B. Z. Ding et al. Fl. Henan 1: 478. 1981; S. Y. He, Fl. Beijing, rev. ed., 1: 244. 1984; J. W. Wang in Fl. Hebei. 1: 466. 1986; Wang, Ziman & Dutton in Fl. China 6: 321. 2001. —A. cathayensis f. hispida (Tamura) Kitag. Neo-Lineam. Fl. Mansh. 290. 1979. Type 热河省: Koryu, 1933-08, T. Nakai, M. Honda & M. Kitagawa s. n. (holotype, TI, non vidi)。朝鲜:Prov. Kannan, Shokori, 1933-07, G. Koidzumi s. n. (paratype, KYO, non vidi)。
  分布于中国河北西部和北部、河南北部,以及朝鲜。生于山地草坡,海拔1 000~2 000 m。
  标本登录 河北:涞源,白石山,刘继孟1628;涞水,刘继孟2190;小五台山,王启无61341;涿鹿,杨朝广826;雾灵山,alt.1 500 m,刘慎谔,傅沛云等4776;东陵,唐进1768;承德,南开大学队127。北京:西灵山,杨朝广1531。   45. 小五台银莲花 图3
  Anemone xiaowutaishanica  W. T. Wang & Bing Liu in Acta Phytotax. Sin. 46(5): 739, fig. 1. 2008; J. C. Zhao et al. Fl. Xiaowutaishan 1:319, fig. 291. 2011; W. T. Wang in High. Pl. China in Colour 3: 405. 2016. Type 河北:小五台山,山涧口,alt.1 692 m,山谷石边草丛,萼片白色,背面淡紫色,2007-09-15,刘冰983 (holotype, PE);同地,2006-09-03,刘冰415 (paratype, PE)。
  特产于中国河北小五台山。生于山谷石边草丛,海拔1 692 m。
  46. 宝兴银莲花 图版7: 5-8
  Anemone baoxingensis W. T. Wang, sp. nov. Holotype 四川 (Sichuan):寶兴,邓池沟 (Baoxing Xian, Dengchigou), alt. 3 360 m,林下,花白色 (in forest, fls. white), 1933-06-26,杜大华 (D. H. Du) 4305 (PE)。
  Perennial Herb ca. 28 cm tall. Rhizome vertical, ca. 1.8 cm long, 1 cm across. Basal leaves ca. 5, long petiolate; blades papery, cordate-pentagonal, 5-8.5 × 9-10 cm, 3-sect, central segment broadly rhombic, 4.5-6 cm broad,3-cleft, lobes 3-lobed and with lobules narrowly ovate and 1-2-dentate or entire, lateral segments obliquely and broadly rhombic, unequally 2-parted, with upper lobes somewhat similar to central segment; surfaces adaxially appressed-pubescent, abaxially on nerves sparsely pubescent; petioles14-16 cm long, at base narrowly vaginate, with scapes spreading-villous (hairs 3-5 mm long). Scapes ca. 3, 21-27 cm long; cymes umbelliform, 3-flowered; involucral bracts 3, sessile, unequal, broadly rhombic, 4.5-5.5 × 4-5.5 cm, 3-parted, somewhat similar to the central segment of basal leaf, rarely rhombic or narrow-rhombic, 1-2.5 cm broad; pedicels slender, 3.5-4.5 cm long, withsparse hairs. Flower: Sepals 5, white, spatulate, 14-18 × 5-6.5 mm, at apex acute, at base attenuate, glabrous, 5-nerved. Stamens ca. 30, glabrous, filaments linear, rarely lanceolate-linear, 1.2-3 mm long; anthers dark-blue, broadly oblong, 1 × 0.4-0.6 mm, apex obtuse. Carpels 6-9, glabrous; ovaries compressed, elliptic, 2.2 × 1 mm; styles 0.8-1 mm long, straight or slightly recurved; stigmas linear, ca. 0.8 mm long.
  特产于中国四川宝兴。生于山谷林下,海拔3 360 m。
  47. 腾冲银莲花 图4
  Anemone tengchongensis  W. T. Wang in Bull. Bot. Res. Harbin 167(2): 160, fig. 1: 5-8. 1996. Holotype 云南:腾冲,狼牙山,alt.3 900~4 000 m,山坡竹林,花白色,1964-06-07,武素功7070 (KUN)。
  A. narcissiflora L. ssp. crinita auct. non (Juz.) Kitag.: Ziman in Fl. China 6: 321. 2001.
  A. narcissiflora auct. non L. : Ziman et al. in J. Jap. Bot. 80(5): 284. 2005.
  特产于中国云南腾冲。生于山地竹林中,海拔3 900~4 000 m。
  48.  垂花银莲花 图版19: 1-2
  Anemone nutantiflora W . T. Wang & L.
  Q. Li in Acta Bot. Yunnan. 8(3): 265, fig. 2: 1-2. 1986; W. T. Wang in Fl. Yunnan 11: 203. 2000. Type 云南:泸水,片马丫口,alt. 3 200 m,竹林下,1978-08-03,刘伦辉s. n. (holotype, KUN; isotype, PE)。   特产于中国云南泸水。生于山地竹林中,海拔3 200 m。
  49. 迭裂銀莲花 图版17: 1-3
  Anemone imbricata  Maxim. Fl. Tangut. 8, t. 22: 1-6. 1889; Hemsl. & Pearson in J. Linn. Soc. Bot. 35: 162. 1902; Finet & Gagnep. in Bull. Soc. Bot. France 51: 73. 1904; Ulbr. in Bot. Jahrb. 37: 243. 1905; Hand. -Mazz. in Acta Hort. Gotob. 13: 178. 1939; Lauener in Not. Bot. Gard. Edinb. 23: 193. 1959; Iconogr. Corm. Sin. 1: 731, fig. 1462. 1972; W. T. Wang in Fl. Reip. Pop. Sin. 28: 55, pl. 14: 1-3. 1980; et in Fl. Xizang. 2: 83. 1985; L. H. Zhou in Fl. Qinghai. 1: 341, pl. 74: 13-15. 1997; W. T. Wang in High. Pl. Chian 3: 498, fig. 798. 2000; Wang, Ziman & Dutton in Fl. China 6: 322. 2001; Ziman et al. in Edinb. J. Bot. 64(1): 90. 2007; et in J. Jap. Bot. 83(3): 150. 2008. —A. obtusiloba D. Don ssp. imbricata (Maxim.) Bruhl in Ann. Bot. Gard. Culc. 5: 78. 1896.  —Anemonastrum imbricatum (Maxim.) Holub in Folia Geobot. Phytotax. 8: 158. 1973. Type 西藏北部:Yangtze河,1884-06,N. M. Przewalski s. n. (holotype, LE; isotype, PE)。
  分布于中国四川西部(海拔4 000~4 700 m)、甘肃(2 500~3 800 m)、青海南部和东部(3 800~4 800 m)、西藏(3 800~5 300 m)。生于山坡草地、灌丛、多石山坡或林中。
  标本登录 四川:乡城,alt. 4 700 m,青藏队3905;雅江,姜恕5083;康定,折多山,alt.4 300 m,杨竟生91-117;大炮山,alt. 4 700 m,Harry Smith 11512;金川,李馨77951;壤塘,姜恕9029;德格,崔友文4960;石渠,alt. 4 000 m,姜恕9249。甘肃:迭部,alt. 2 500 m,白龙江队1868;天祝,alt. 3 200 m,河西队236;肃南,河西队350。青海:可可西里,alt. 4 800 m,武素功692A (KUN);治多,alt. 4 300 m,周立华51;玉树,何廷农,B. Bartholomew等2059;玛沁,何廷农688;同德,吴玉虎5518;大通,alt. 3 800 m,杨亲二95-35;祁连,青甘队2811。西藏:江达,alt. 3 800 m,青藏队33;昌都,青藏队3609;左贡。alt.5 100~5 300 m, D. E. Boufford等29459;索县,王金亭3300;比如,陶德定11380,安多,alt.4 650 m,王金亭3722;帕里,alt. 5 300 m,陶德定10807;班戈,郎楷永9428;土门格拉,杨金祥1939;吉隆,alt. 4 900 m,青藏队植被组5050。
  50. 红萼银莲花
  Anemone smithiana  Lauener & Panigrahi in Not. Bot. Gard. Edinb. 33: 491. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 53. 1980; et in Fl. Xizang. 2: 83. 1985; Wang, Ziman & Dutton in Fl. China 6: 322. 2001; Ziman et al. in J. Jap. Bot. 81(1): 5. 2006,et 83(3): 151. 2008. Holotype Nepal: Telley, alt.15 000 ft., 1930, Lall Dhwoj 460 (E; photo, PE).
  分布于中国西藏南部,以及不丹和尼泊尔。生于多石山坡或灌丛中,海拔3 800~4 150 m。
  标本登录 西藏:吉隆,西藏中草药队595,693;聂拉木,青藏队5749,青藏队植被组4246。
  51. 多裂银莲花 图版19: 3-6
  Anemone multilobulata  W. T. Wang & L. Q. Li in Acta Bot. Yunnan. 8(3): 264, fig. 2: 3-6. 1986; W. T. Wang in Fl. Yunnan 11: 204. 2000. Type 云南:碧江,高黎贡山,28分界碑,alt. 3 600 m,花黄色,1978-07-14,碧江队1134(holotype, KUN);同地,渣拉河左侧主峰,1978-07-13,碧江队10800 (paratype, KUN); 同地,1978-07-12,刘伦辉s. n. (paratype, KUN)。   A. narcissiflora L. ssp. protracta auct. non (Ulbr.) Ziman & Pedor: Ziman in Fl. china 6: 321. 2001.
  A. narcissiflora auct. non L.: Ziman et al. in J. Jap. Bot. 80(5): 284. 2005.
  特产于中国云南福贡和碧江高黎贡山。生于海拔3 600~3 800 m一带山地。
  标本登录 云南:福贡,高黎贡山,1978-07-12,?s. n.; 碧江,海拔3 800 m,1978-07-13, ? s. n.
  系2. 复伞银莲花系
  Ser. Involucellatae  Ulbr. in Bot. Jahrb. 37: 209, 267. 1906; Tamura in Sci. Rep.Osaka Univ. 16: 22. 1961; Ziman et al. in J. Jap. Bot. 80(5): 289 2005. Type: A. polyanthes D. Don.
  伞状复伞花序复杂,具较多花。
  4种,分布于喜马拉雅山区、中国华北东部以及日本,在中国均有分布。
  52. 复伞银莲花
  Anemone tetrasepala Royle, Ill. Bot. Himal. 53: 1839; Hook. f. & Thoms. in Hook. f. Fl. Brit Ind. 1: 10. 1872; Ulbr. in Bot. Jahrb. 37: 268. 1906; W. T. Wang in Fl. Reip. Pop. Sin. 28: 55. 1980; et in Fl. Xizang. 2: 84 1985; Wang, Ziman & Dutton in Fl. China 6: 319. 2001; Ziman et al in J. Jap. Bot. 81(1) 10. 2006, et 83(3): 151. 2008. HolotypeKashmir: Jum Boo. 1828, J. F. Royle s. n. (non vidi).
  分布于中國西藏南部,以及克什米尔、巴基斯坦和阿富汗。生于山地林间草地或林下,海拔2 900~3 100 m。
  标本登录 西藏:吉隆,青藏队7013;聂拉木,樟木,张永田,郎楷永4501。
  53.  山东银莲花 图版17: 7-9
  Anemone sikokiana  (Makino) Makino in Bot. Mag. Takyo 27: 116. 1913; Tamura & Shimizu in Satake et al. Wild Flow. Japan 2: 68. 1982; Ziman et al. in J. Jap. Bot. 81(1): 13. 2006, et 83(3): 151. 2008; W. T. Wang in High. Pl. China in Colour 3: 403. 2016; L. Xie in Medic. Fl. China 3: 336. 2016. — A. narcissiflora L. var. shikokiana Makino (in Bot. Mag. Tokyo 8: 172. 1894, nom. nud. ) in Bot. Mag. Tokyo 16: 58. 1902. — Anemonastrum sikokianum (Makino) Holub in Folia Geobot. Phytotax. 8: 58. 1973. Syntype 日本:Prov. Iyo, Mt. Ishizuchi, 1891-08, L. Doi s. n. (RI, non vidi).
  A. chosenicola Ohwi in Acta Phytotax. Geobot. 7: 47. 1938.
  A. schantungensis Hand. -Mazz. in Acta Hort. Gotob. 13: 181. 1939; W. T. Wang in Acta Phytotax. Sin. 12(2): 112. 1974. — A. chosenicola Ohwi var. schantungensis (Hand.-Mazz.) Tamura in Acta Phytotax. Geobot. 16: 110. 1958; W. T. Wang in Fl. Reip. Pop. Sin. 28: 55, pl. 14: 7-9. 1980.
  分布于中国山东东部、河北北部以及日本。生于山地草丛,海拔600~1 100 m。
  标本登录 山东:昆嵛山,alt. 900~960 m,刘慎谔,刘继孟1489,刘继孟550,PE考察队56-2994,刘长江141;青岛,白云洞,F. H. Sha 544;崂山,焦启元2768,P. Licent 13390,周太炎1290。河北:小五台山,?7118。
  54. 柄萼银莲花
  Anemone elongata  D. Don, Prodr. Fl. Nepal. 194. 1825; Hook f. & Thoms. in Hook. f. Fl. Brit. Ind. 1: 10. 1872; Ulbr. in Bot. Jahrb. 37: 268. 1906; Wang, Ziman & Dutten in Fl. China 6: 320. 2001; Ziman et al. in J. Jap. Bot. 81(1): 12. 2006, et 83(3): 151. 2008. — Anemonastrum elongatum (D. Don) Holub in Folia Geobot. Phytotax. 8: 165. 1973. Holotype: Nepal, 1821, Wallich 4692 (non vidi; photo, PE).   分布于中国西藏,以及尼泊尔、印度西北部 (fide Ziman et al., 2006)。
  标本登录 西藏:Lende Khola, alt.9 500 ft., 1939-07-11, Polunin 861 (BM, non vidi, fide Ziman et al., 2006)。
  55. 多花银莲花
  Anemone polyanthes  D. Don, Prodr. Fl. Nepal. 194. 1825; Hook. f. & Thoms. in Hook. f. Fl. Brit. Ind. 1: 9. 1872; Franch. in Bull. Soc. Bot. France 33: 367. 1886; Forbes & Hemsl. in J. Linn. Soc. Bot. 23: 12. 1886; Ulbr. in Bot. Jahrb. 37: 267. 1906; Ziman et al. in J. Jap. Bot. 81(1): 9. 2006, et 83(3): 151. 2008. — A. narcissiflora L. var. polyyanthes (D. Don) Finet & Gagnep. in Bull. Soc. Bot. France 51: 56. 1904. — Anemonastum polyanthes (D. Don) Holub in Folia Geobot. Phytotax. 8: 165. 1973. Holotype: Himalaya, Wallich 4691b (non vidi).
  分布于中国云南西北部(洱源,fide A. Franchet, 1886, et Ziman et al., 2006),以及巴基斯坦、尼泊尔、不丹、印度北部和缅甸。
  标本登录 云南:洱源,Mt. Hee-chan-men prope Kan-hay-tze, alt. 2 800 m, 1884-05, J. M. Delavay 1110 (photo, PE); 同地,Mt. Hee-chan-men, Hyra Yangkong, 1887-05-06, J. M. Delavay 110 (K, fide Ziman et al., 2006)。
  組5.  短柱银莲花组
  Sect.Brachycladus W. T. Wang, sect. nov. Type: A. brevistyla Chang ex W. T. Wang
  Ob habitus, sepala 5 vel 6 alba, staminum filamenta filiformia et pollinis grana 6-7-colpata sectio nova haec est arcte affinis Sect. Stoloniferae (Ulbr.) Juz., a qua carpellorum stylis conspicuis rectis ovariis leviter brevioribus, stigmatibus inconspicuis vel conspicuis minutis differt.
  Rhizomes subhorizontal or vertical. Basal leaves 2-4, long petiolate, simple and 3-sect or ternate. Cymes 1-flowered; involucral bracts three or two, sessile, 3-parted. Sepals 5 or 6, white. Stamen filaments filiform. Carpels with ovaries densely pubescent and straight styles slightly shorter than ovaries and glabrous; stigmas inconspicuous or conspicuous, but small.
  Two species, distributed in western Sichuan Province.
  根状茎近水平或垂直。基生叶2~4,具长柄,为单叶,3全裂或为三出复叶。聚伞花序有1花;总苞苞片3或2,无柄,3深裂。萼片5或6,白色。雄蕊花丝丝形;花粉具6~7条沟(贡嘎山的花粉尚不了解)。心皮的子房密被柔毛,花柱直,比子房稍短,无毛;柱头不明显或明显,小。
  2种,分布于四川西部。
  56. 短柱银莲花 图版20: 4-6
  Anemone brevistyla Chang ex W. T. Wang in Acta. Phytotax. Sin. 12(2): 162, pl. 2: 4-6. 1974; et in Fl. Reip. Pop. Sin. 28: 16, pl. 3: 4-6. 1980; Wang, Ziman & Dutton in Fl. China 6: 318. 2001. Holotype 四川:天全,1936,曲桂岭2119 (PE)。
  A. baicalensis auct. non Turcz.: Ziman et al. in J. Jap. Bot. 79(5): 296. 2004.
  特产于中国四川西部(天全、泸定、都江堰)。生于山地林边,海拔约2 200 m。
  标本登录 四川:泸定,磨西镇,alt. 2 280 m,成都生物所植被组30284;都江堰,?960073。   57.  贡嘎银莲花  图版21: 11-13
  Anemone gonggaensis W. T. Wang, sp. nov. Holotype 四川(Sichuan): 康定,贡嘎山,贡嘎寺 (Kangding, Gongga Shan, Gongga Temple), alt. 3 720 m,林缘,花白色 (forest margin, fls. white),2018-07-18,杨永,李海文,杨智,王宇 (Y. Yang, H. W. Li, Z. Yang & Y. Wang) GGS316 (PE)。
  Perennial herbs 16-19 cm tall. Rhizomes vertical, ca. 1.2 cm long, at apex 5 mm across. Basal leaves 3-4, ternate, long petiolate; blades 2.5-5.5 × 2.5-6.5 cm; cental leaflet with petiolule 1-10 mm long, broadly rhombic, 2-3 cm broad, 3-parted slightly exceeding the middle of leaflet, central primary lobe rhombic, at apex acute, 3-lobed, central secondary lobe also rhombic and 3-lobed, lateral primary lobes oblique, 2-5-denticulate, lateral leaflets with petiolules 1-5 mm long or sessile, obliquely rhombic, 2-3.6 × 1.4-3 cm, unequally 3-parted; surfaces adaxially and abaxially glabrous or subglabrous; petioles 5-15 cm long, on apex and base with a few hairs, elsewhere glabrous. Scapes solitary, 5.5-12.5 cm long, only on apex with a few hairs; cymes1-flowered; involucral bracts 2, sessile, glabrous, unequal, the larger one broadly rhombic, 3.2-5 × 3-5 cm, 3-parted, and the smaller one rhombic-ovate, 3-4.3 × 2.3 cm, also 3-parted; pedicels slender, 4-6 cm long, appressed-pubescent. Flower: Sepals 6, white, broadly elliptic or orbicular-elliptic, 20-25 × 11-15 mm, glabrous, apex rounded or obtuse. Stamens ca. 100, glabrous; filaments 4-5 mm long, filiform, but near base slightly dilated; anthers yellowish, broadly elliptic, 0.8-1 × 0.5-0.8 mm, apex obtuse. Carpels 20-30; ovaries narrowly ovoid, 1 mm long, densely appressed-puberulous; styles0.6-0.8 mm long, glabrous; stigmas situated on apex of style, ellipsoid, 0.3-0.4 mm long.
  特产于中国四川省康定市贡嘎山。生于山地林缘,海拔3 721 m。
  组6.  鹅掌草组
  Sect. Stolonifera (Ulbr.) Juz. in Fl. URSS 7: 252. 1937; W. T. Wang in Fl. Reip. Pop. Sin. 28: 15. 1980; Tamura in Acta Phytotax. Geobot. 42: 179. 1991; et in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17aIV: 341. 1995; Ziman et al. in J. Jap. Bot. 79(1): 49, (5): 294. 2004. —Subsect. Stolonifera Ulbr. in Bot. Jahrb. 37: 195, 230. 1905. Lectotype: A. baicalensis Turcz.
  Sect. Deinostigma W. T. Wang in Guihaia39(1): 7. 2019. Type: A. weixiensis W. T. Wang.
  根狀茎短或长。基生叶1~3(~7),稀较多,3全裂,稀3半裂。聚伞花序有1~3(~6)花;总苞苞片3,稀2,无柄,3深裂或3浅裂,稀不分裂。萼片5。雄蕊花丝丝形;花粉具4~10条沟。心皮的子房被柔毛,稀无毛;花柱极短或不存在;柱头明显,球形,卵球形或陀螺形,稀长圆形。染色体基数x=7。
  约15种,分布于亚洲东部温带和暖温带地区。中国有10种,广布于西南、长江流域、秦岭山区和东北诸省,多数种(6种)分布于西南山区;可划分为4系。   系1.  鹅掌草系
  Ser. Flaccidae Juz. in Fl. URSS 7: 252. 1937; Ziman et al in J. Jap. Bot. 79(1): 49 et (5): 297. 2004. Type: A. flaccida F. Schmidt.
  根状茎不生出匍匐茎。基生叶1~3(~7),3全裂。聚伞花序具1~5花。萼片5(~8)。
  约6种。
  59. 鹅掌草 图版20: 1-3
  Anemone flaccida F. Schmidt in Mem. Acad. Sci. Petersb., ser. 7, no. 2: 103. 1868; Franch. in Bull. Soc. Bot. France 33: 366. 1886; Forbes & Hemsl. in J. Linn. Soc. Bot. 23: 11. 1886; Finet & Gagnep. in Bull. Soc. Bot. France 51: 70. 1904; Pei in Contr. Biol. Lab. Sci. Soc. China, Bot. ser., 9(1): 3. 1933; Iconogr. Corm. Sin. 1: 726, fig. 1451. 1972; Fl. Tsinling. 1(2): 280. 1974; W. T. Wang in Fl. Reip. Pop. Sin. 28: 16, pl. 3: 1-3. 1980; B. Z. Ding et al. Fl. Henan. 1: 478. 1981; X. W. Wang in Fl. Anhui. 2: 327, fig. 632. 1987; Y. K. Li in Fl. Guizhou. 3: 80, pl. 32: 4-7. 1990; Z. H. Lin in Fl Zhejiang. 2: 279, fig. 2-371. 1992; W. T. Wang in Fl. Yunnan. 11: 187. 2000; et in High. Pl. China 3: 491, fig. 783. 2000; K. M. Liu, Fl. Hunan. 2: 658, fig. 2-515. 2000; S. H. Fu, Fl. Hupeh. 1: 351, fig. 490. 2001; Wang, Ziman & Dutton in Fl. China 6: 310. 2001; W. T. Wang in Fl. Jiangxi 2: 159. 2004; Ziman et al. in J. Jap. Bot. 79(5): 277. 2004, et 83(3): 147. 2008; M. B. Deng & K. Ye in Fl. Jiangsu 2: 87, fig. 2: 140. 2013; L. Xie in Medic. Fl. China 3: 312. 2016. —Anemonoides flaccida (F. Schmidt) Holub in Folia Geobot. Phytotax. 8: 166. 1973. — Arsenjevia flaccida (F. Schmidt) Starod. Vetrenytry 123. 1991.
  — Anemonastrum flaccidum (Fr. Schmidt) Mosyakin in Phytoneuron 55: 5. 2018. Holotype俄罗斯(俄国)“Due and the village of Mgachi in Sakhalin, Augustinowicz s. n.”(LE, non vidi)。
  A. baicalensis Turcz. var. laevigata Gray in Perry, Narret. Exp. Ch. Jap. 2: 306. 1857; Hand. -Mazz. in Acta Hort. Gotob. 13: 175. 1939. — A. laevigata (Gray) Koidz. in Bot. Mag. Tokyo 43: 395. 1929.
  58a. var. flaccida
  分布于中国云南西北(海拔2 800~3 600 m)、四川(1 500~2 500 m)、贵州(650~1 800 m)、湖北西部、甘肃南部(1 600~2 700 m)、陕西南部(1 100~2 200 m)、湖南(1 000~1 200 m)、江西、安徽南部(930~1 500 m)、江蘇南部和浙江西北部(800 m),以及俄罗斯(俄国)远东地区和日本。生于山地林下、草坡或灌丛中。
  本种的根状茎可供药用,有祛风除湿,强壮筋骨之效,用于跌打损伤、风湿痛等症(谢磊等,2016)。
  标本登录 云南:维西,王启无63633,冯国楣3814,杨亲二,孔宏智98-437;丽江,秦仁昌20961;鹤庆,秦仁昌23465;兰坪,蔡希陶53799;大理,杨亲二93-70。四川:峨眉山,关克俭373;宝兴,杜大华4867;崇州,冯正波4352;都江堰,冯正波2007-4;安县,朱大海3733;平武,川大生物系10839。重庆:金佛山,李国凤61060;武隆,植物资源队59-148;彭水,植物资源队59-2904;奉节,周洪富110986;巫山,杨光辉57987。贵州:遵义,黔北队284;梵净山,张志松等401686;凯里,黔南队1105。湖北:利川,罗毅波,向巧苹202;宜昌,龙茹09-42。甘肃:夏河,王作宾1173;舟曲,姜恕407;成县,王庆瑞11526;天水,王庆瑞11360。陕西:凤县,罗毅波10;佛坪,郭本兆1411;太白山,alt.1 100~1 300 m,王作宾6510,傅坤俊2536。湖南:新宁,罗仲春1767;衡山,李明江739。江西:安福,武功山,1929-04-10,钟观光s. n. 安徽:金寨,邓懋彬,郎楷永93-005;岳西,资源植物队59-46;绩溪,黄成林932。江苏:镇江,H. Migo s. n.;苏州,凌苹苹,孙三省66。浙江:天目山,贺贤育20942;安吉,俞立鹏,邓懋彬97-69。   58b. 展毛鹅掌草 (变种)
  var.hirtella W. T. Wang in Fl. Reip. Pop. Sin. 28: 18, 349. 1980; Wang, Ziman & Dutton in Fl. China 6: 311. 2001; Ziman et al. in J. Jap. Bot. 79(5): 298. 2004. Holotype 湖北:利川,alt. 1 000 m,溪边,1946-09-04,郑万钧,华敬灿871 (PE)。
  特产于中国湖北利川。生于山地溪边,海拔1 000 m。
  59. 细茎银莲花 图版28: 5
  Anemone rossii S. Moore in J. Linn. Soc. Bot. 17: 376, t. 16: 1-2. 1879; Forbes & Hemsl. In l. c. 23: 11. 1886; Kom. in Acta. Hort. Petrop. 22: 264. 1902; Fl. Pl. Herb. Chinae Bor.-Or. 3: 154. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 18, pl. 16: 5. 1980; S. H. Li, Fl. Liaoning 1: 511, pl. 211: 3-4. 1988; Y. Zhou, Colour. Ill. Pl. Wet Area. NE China 347. 2019. — A. baicalensis ssp. glabrata var. rossii (S. Moore) Kitag. in Rep. Inst. Sci. Res. Manch. 4(7): 81. 1940; Wang, Ziman & Dutton in Fl. China 6: 312. 2001. — Anemonoides rossii (S. Moore) Holub in Folia Geobot. Phytotax. 8: 155. 1973. —Arsenjevia rossii (S. Moore) Starod. Vetrenytsy 122. 1991. Described from Fengcheng, Liaoning Province.
  A. baicalensis auct. non Turcz.: Ziman et al. J. Jap. Bot. 79(5): 294. 2004.
  分布于中国辽宁(凤城)、吉林东部以及朝鲜北部。生于山地草坡或林下。
  标本登录 吉林:安图,长白山,1958-05-04,?4128;长白山,方振富等1289,1350;桦甸,钓鱼台,马XX323 (IFP)。
  60.  维西银莲花 图版22
  Anemone weixiensis  W. T. Wang in Guihaia 39(1): 9, fig. 1. 2019. Holotype 云南:维西,栗地坪,alt. 3 100 m,林间草地,花白色,2008-07-28,杨亲二,袁琼2042 (PE)。
  特产于中国云南维西。生于山地林间草地,海拔3 100 m。
  61.  芹叶银莲花 图版21: 1-3
  Anemone saniculiformis C. Y. Wu ex W. T. Wang in Acta Phytotax. Sin. 12(2): 164, pl. 8: 7. 1974; et in Fl Reip. Pop. Sin. 28: 19. 1980. — A. baicalensis Turcz. var. saniculiformis (C. Y. Wu ex W. T. Wang) Ziman & Dutton in Fl. China 6: 312. 2001; Ziman et al. in J. Jap. Bot. 79 (5): 296. 2004. Type: 寶兴,1954,宋滋圃38347 (holotype, KUN);同地,蒲溪沟,alt. 2 300 m, 1959-05-02,四川经济植物队59-245 (paratype, NAS)。
  特产于中国四川宝兴。生于山地路边林中,海拔2 300~2 400 m。
  标本登录 四川:宝兴,小灯笼沟,alt. 2 400 m,1958-06,张秀实,任有铣5305。
  62. 滇川银莲花  图版23: 10-12
  Anemone delavayi Franch. in Bull. Soc. Bot. France 33: 366. 1886; Pl. Delav. 9. 1889; Finet & Gagnep. in Bull. Soc. Bot. France 51: 66. 1904; Ulbr. in Bot. Jahrb. 37: 233. 1905; Hand. -Mazz. in Acta Hort. Gotob. 13: 175. 1939; Iconogr. Corm. Sin. 1: 726, fig. 1452. 1972; W. T. Wang in Fl. Reip. Pop. Sin. 28: 22, pl. 4: 10-12. 1980; et in Fl. Yunnan. 11: 188, pl. 55: 7-9. 2000; Wang, Ziman & Dutton in Fl. China 6: 311. 2001; Ziman et al. in J. Jab. Bot. 79(5): 299. 2004, et 83(3): 147. 2008; L. Xie in Medic. Fl. China 3: 316. 2016. Holotype 云南:Mao-kon-tchang prope Ta-pin-tze, alt.2 200 m, 1885-05-18, J. M. Delavay 1504 (P, non vidi; photo, PE)。   A. oligocarpa Pei in Contr. Biol. Lab. Sci. Soc. China 9: 3, fig. 1. 1933. —A. delavayi var. oligocarpa (Pei) Ziman & Dutton in Fl. China 6: 311. 2001; Ziman et al. in J. Jap. Bot. 79(5): 299. 2004. Holotype 四川:九龙,alt. 3 000 m,山谷阴地,1939-05-23,郑万钧988 (non vidi)。
  分布于中国云南西北部、西藏东南部(察隅)和四川西南部(九龙)。生于山地林下、草坡或溪边,海拔2 200~3 000 m。
  标本登录 云南:贡山,昆明工作站9335;怒江,澜沧江分水岭,俞德浚19076;怒江,俅江分水岭,俞德浚19291;中甸,alt. 2 400~2 700 m,俞德浚11411,11557;维西,王启无64559,67814; 丽江,王启无71390, 秦仁昌20538。西藏:察瓦龙,松塔雪山,alt. 3 000 m,青藏队7742。
  63. 福贡银莲花 图版21: 4-7
  Anemone fugongensis W. T. Wang, sp. nov. Type 云南 (Yunnan):福贡县,匹河 (Fugong Xian, Pihe), alt. 2 910 m,林中,花白色 (in forest, fls. white), 2008-05-15,金效华,张亭 (X. H. Jin & T. Zhang) 61(holotype, PE);福贡县,鹿马登 (Fugong Xian, Lumadeng), alt.2 078 m, 2008-05-14,金效华,张亭46 (paratype, PE)。
  63a. var. fugongensis
  Small perennial herbs 5-9.5 cm tall. Rhizomes vertical, ca. 5 × 3 mm. Basal leaves 1-2, long petiolate; blades thinly papery, cordate-pentagonal, 1.7-3.2 × 2.5-3.7 cm, 3-sect, central segment broad-rhombic, 1.5-2 cm broad, at apex acute, below the middle with entire margins, above 3-lobulate, with lobes 2-4-denticulate, lateral segments broadly obovate or flabellate, as long as or longer and larger than central segment, 2-lobulate or 2-cleft, secondary lobes incised-dentate; surfaces adaxially and abaxially appressed-pubescent; petioles 2-5.8 cm long, with scapes sparsely pubescent. Scapes 1-2, 3.5-7 cm long; cymes 1-flowerd; involucral bracts 3, sessile, broad-rhombic, 5-8 × 5-15 mm,3-lobed, 3-cleft or 3-parted, with lobes ovate, triangular or linear, entire, rarely 1-2-denticulate, sparsely pubescent; pedicels 1.5-3.5 cm long, sparsely pubescent, rarely near apex with dense hairs. Flower: Sepals 5, white, obovate, 5.5-10 × 4-7 mm, on apex rounded, adaxially glabrous, abaxially with sparse hairs, 5-nerved. Stamens 20-25, glabrous; filaments filiform, 2-4 mm long; anthers oblong, ca. 1 mm long. Carpels 5-8; ovaries narrow-ovoid, 1.5 × 0.7 mm, densely appressed-pubescent; styles ca. 0.5 mm long; stigmas oblong, 0.6-1 mm long.
  產于中国云南福贡。生于山地林中或林边,海拔2 078~2 910 m。
  63b. 泸水银莲花 (变种) 图版28: 6-7
  var. lushuiensis  W. T. Wang, var. nov. Type
  云南 (Yunnan):泸水县,高黎贡山 (Lushui Xian, Gaoligong Shan), alt. 3 000 m,山坡,花淡紫色 (on slope, fls. purplish), 1981-06-03,赵魁义 (K. Y. Zhao) 148 (holotype, PE);泸水,自片马至竹节湖途中 (Lushui, on the way from Pianma to Zhujie Lake), alt. 3 500 m,山坡竹林中,花淡蓝紫色 (in bamboo thicket on slope, fls. bluish-purplish), 1981-05-31,赵魁义103 (PE)。   A. var. fugongensi differt sepalis purpurascentibus, ovariis glabris.
  产于中国云南省泸水县。生于山地山坡上或竹林中,海拔3 000~3 500 m。
  系2.  毛果银莲花系
  Ser. Baicalenses Juz. in Fl. URSS 7: 250. 1937.
  — Ser. Stoloniferae. Ziman et al. J. Jap. Bot. 79(1): 49. 2004. — Ser. Stolonifera. Ziman, Kadota & Bulakh in J. Jap. Bot. 79(5): 294. 2004. Type:  A. baicalensis Turcz.
  根状茎生出一条匍匐茎。基生叶3全裂或3半裂。聚伞花序有1~2花。萼片5~8。
  3种。
  64. 毛果银莲花 图版23: 1-6
  Anemone baicalensis Turcz. in Bull. Soc. Nat. Mosc. 15: 42. 1842; Maxim. in Acta Hort. Perop. 11. 22. 1890; Kom. in l. c. 22: 261. 1903; Finet & Gagnep. in Bull. Soc. Bot. France 51: 70. 1904; Ulbr. in Bot. Jahrb. 37: 230. 1905; Pei in Contr. Biol. Lab. Sci. Soc. China, Bot. ser., 9(1): 2. 1933; Hand. -Mazz. in Acta Hort. Gotob. 13: 174. 1939; Fl. Pl. Herb. Chinae Bor. -Or. 3: 154. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 20, pl. 4: 1-6. 1980; S. H. Li, Fl. Liaoning. 1: 513, pl. 211:  1-2. 1988; W. T. Wang in High. Pl. China 3: 492, fig. 784. 2000; Wang, Ziman & Dutton in Fl. China 6: 311. 2001, p. p. excl. syn. A. baicalensis var. glabrata Maxim.; Ziman et al. in J. Jap. Bot. 79(5): 294. 2004, p. p. excl. syn. A. rossii S. Moore et A. brevistyla Chang ex W. T. Wang, et 83(3): 147. 2008; X. D. Li, Y. Y. Zan & S. Huang in Fl. Shennongjia 1: 283. 2017. —Anemonoides baicalensis (Turcz.) Holub in Folia Geobot. Phytotax. 8: 115. 1973. — Aresenjevia baicalensis (Turcz.) Starod. Vetrenytsy 122. 1991.  — Anemonastrum baicalense (Turcz.) Mosyakin in Phytoneuron 55: 8. 2018. Discribed from central Siberia.
  A. ulbrischiana Diels ex Ulbr. in Notizbl. Bot. Gart. Berl. 10: 876. 1929; Fl. Tsinling1(2): 281, fig. 241. 1974.
  64a. var. baicalensis
  分布于中国四川北部、重庆北部、湖北西部、甘肃南部、陕西南部、河北北部、辽宁、吉林和黑龙江,以及朝鲜和俄罗斯西伯利亚地区。生于山地林下、林边或草坡上,海拔500~3 100 m。
  标本登录
  四川: 理县,何铸12617(SZ);阿坝,alt. 2 965 m, D. E. Boufford等43645。甘肃:文县,alt. 2 170~2 425 m, D. E. Boufford等37815;舟曲,alt. 3 100 m,王作宾14437;夏河,王作宾7260。陕西:太白山,alt. 2 800 m,杨金祥858,秦岭队10257。河北:?,1929-07-06,P. Licent. 6688。辽宁:草河口,赵大昌等602。吉林:静宁县,Tang Ling-yu 12979;长白山,alt. 910 m,森林组045;抚松,通化组236;威虎岭,孔宪武1539。黑龙江:小兴安岭,宋朝枢90。
  64b.  甘肃银莲花 (变种)
  var. kansuensis (W. T. Wang) W. T. Wang in Fl. Reip. Pop. Sin. 28: 20. 1980; Wang, Ziman & Dutton in Fl. China 6: 312. 2001.
  — A. kansuensis W. T. Wang in Acta Phytotax. Sin. 12(2): 163, pl. 8: 6. 1974. Type 甘肅:武都,alt.2 500 m,1930-06-29,郝景盛432 (holotype, PE);同地,麻柳片,1959-05-14,张志英1447 (paratype, WUK)。   特产于中国甘肃南部(文县、武都)。生于山地林下、草丛或沟边,海拔1 600~2 600 m。
  标本登录 甘肃:文县。alt. 1 600~2 600 m,白水江队0179,3812,5240,5312,5487,5765。
  64c. 光果银莲花 (变种) 图版23: 7-9
  var. glabrata Maxim. Prim Fl. Amur. 18. 1859; Hand. -Mazz. in Acta Hort. Gotob. 13: 175. 1939. p. p. excl. syn. A. rossii S. Moore; Fl. Pl. Herb. Chinae Bor. -Or. 3: 156. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 20. 1980; Ziman et al. in J. Jap. Bot. 79(5): 296. 2004. — A. glabrata (Maxim.) Juz. In Fl. URSS 7: 197. 1937. — A. baicalensis ssp. glabrata (Maxim.) Kitag. in Rep. Inst. Res. Manch. 4: 81. 1940. — Anemonoides glabrata (Maxim.) Starod. Vetrenystsy 122. 1991. Holotype: “Amur River, near the village of Daiso, C. Maximowicz s. n. (LE, nov vidi).
  分布于中国吉林和黑龙江以及俄罗斯远东地区。生于山地云杉林或红松林边。
  标本登录 黑龙江: 伊春,中德队7877,吴征镒128。
  65. 川西银莲花 图版20: 7-10
  Anemone prattii Huth ex Ulbr. in Bot. Jahrb. 36, Beibl. 80: 4, fig. 1-2. 1905 et in l. c. 37: 232. 1905; Hand. -Mazz. in Acta Hort. Gotob. 13: 175. 1939; W. T. Wang in Fl. Reip. Pop. Sin. 28: 18, pl. 3: 7-10. 1980; et in Fl. Yunnan. 11: 188. 2000; Wang, Ziman & Dutton in Fl. China 6: 312. 2001; Ziman et al. in J. Jap. Bot. 79(5): 296. 2004. — Anemonastrum prattii (Huth ex Ulbr.) Mosyakin in Phytoneuron 55: 5. 2018. Holotype 四川:康定,alt. 3 000 m, A. E. Pratt 773 (non vidi)。
  65a. var. Prattii
  分布于中国四川西部和云南北部(茨开,巧家)。生于山地草坡、林边或林下,海拔1 700~3 000 m。
  标本登录 四川:石棉,谢朝俊39734,40035,40059;天全,蒋兴麐33722;二郎山,alt.1 700 m, T. Naito,郎楷永等153;峨眉山,俞德浚361,461,656,杨光辉54210,54266,关克俭440;马边,方文培457;宝兴,alt. 2 300 m,经济植物队59-247;都江堰,alt. 1 740~2 230 m, 汪发瓒20845,杨亲二95-028。
  在上列15号标本中,只有谢朝俊39734和关克俭440二号标本的根状茎均生出一条具1枚小块茎的细长匍匐茎,其他13号标本均无匍匐茎。
  65b. 疏毛川西银莲花 (变种) 图版21:8-10
  var. sparsipilifera, W. T. Wang, var. nov. Holotype 四川 (Sichuan): 泸定县,磨西,小河子沟 (Luding Xian, Mexi, Xiaohezigou), alt. 2 460 m, 路边林下(in forest by road),2018-05-10,贡嘎山采集队 (Gongga Shan Exped.) 0412 (PE)。
  A. var. prattii differt stolone haud tuberifero, ovariis pilis sparsis 0.1 mm longis obtectis.
  特产于中国四川省泸定县磨西镇。生于山地林下,海拔2 460 m。
  65c. 涪陵银莲花 (变种) 图版24
  var. fulingensis  (W. T. Wang & Z. Y. Liu) W. T. Wang, st. nov — A. fulingensis W. T. Wang & Z. Y. Liu in Acta Phytotax. Sin. 45(3): 290. fig. 1. 2007. Type 重慶:涪陵,大木乡,猫儿沟,alt. 1 420 m, 2006-04-22, 刘正宇,任明波2060366 (holotype, PE);同地,2001-05-06,刘正宇2050007 (paratype, PE)。
  特产于中国重庆涪陵。生于海拔1 420 m 一带山地。
  66. 红叶银莲花 图5
  Anemone erythrophylla Finet & Gagnep. in Bull. Soc. Bot. France 53: 125. 1906; W. T. Wang in Fl. Reip. Pop. Sin. 28: 19. 1980; et in Acta Bot. Yunnan. 6(4): 378. 1984; Wang, Ziman & Dutton in Fl. China 6: 311. 2001. — A. wilsonii Hemsl. in Kew Bull. 1906: 149. 1906, non Ulbr., 1905. Holotype 四川:无准确地点,alt. 7~9 000 ft., 1904, E. H. Wilson 3038 (K, photo, PE)。   产于中国四川西部。生于山地阴湿处或石上,海拔1 850~2 700 m。
  标本登录 四川:天全,二郎山,alt. 1 850 m,杨亚滨21839。
  系3.  鹤峰银莲花系
  Ser. Hofengenses W. T. Wang, ser. nov. Type: A. hofengensis W. T. Wang
  A sectionis hujus seriebus ceteris differt antheris apice minute mucronatis. In seriebus ceteris, antherae apice obtusae, haud mucronatae sunt.
  Rhizome not putting forth a stolon. Basal leaves1-4, 3-sect. Cymes 2-6-flowered. Sepals 5. Anthers on apex minutely mucronate. Carpels 9-19; ovaries hairy; styles very short or absent; stigmas turbinate or ovoid.
  本系的花药顶端具直径约0.1 mm的小尖头,据此可与本组其他系区别。
  根状茎不生出匍匐茎。基生叶1~4,3全裂。聚伞花序有2~6花。萼片5。花药顶端具小尖头。心皮9~19;子房有毛;花柱极短或不存在;柱头陀螺形或卵球形。
  2种。
  67.  鹤峰银莲花 图6
  Anemone hofengensis W. T. Wang in Acta
  Phytotax. Sin. 29(5): 163, fig. 3: 6. 1991; K. M. Liu, Fl. Hunan 2: 660. 2000. — A. flaccida F. Schmidt var. hofengensis Wuzhi in S. H. Fu, Fl. Hupeh. 1: 352, fig. 491. 1976, nom seminud; W. T. Wang in Fl. Reip. Pop. Sin. 28: 18. 1980. —A. flaccida var. hofengensis (W. T. Wang) Ziman & Dutton in Fl. China 6: 311. 2001;Ziman et al. in J. Jap. Bot. 79(5): 298. 2004. Holotype
  湖北:鹤峰,小坪,花白色,1958-08-15,李洪钧6122(PE)。Paratypes (均存PE):湖北:鹤峰,芹草坪,李洪钧6804;恩施,枚桥,方明渊24828。湖南:桑植,沙地坪,刘林翰9044;桑植,天平山,李丙贵,万绍宾750001。
  分布于中国湖南西北、湖北西南和重庆东北部。生于山地草坡、沟边或林边,海拔800~1 800 m。
  标本登录 湖南:桑植,八大公山,alt. 1 250 m,北京队2494。湖北:鹤峰,alt. 800 m,彭辅松1077;恩施,alt. 1 600 m,周洪富,粟和毅108025。重庆:奉节,alt. 1 280~1 800 m,方明渊24203,24248,张泽荣25046,25325,周洪富,粟和毅108132,108347。
  68. 安徽银莲花 图7
  Anemone anhuiensis Y. K. Yang, N. Wang & W. C.Ye in J. Wuhan Bot. Res. 7(4): 327, fig. 1. 1989. — A. flaccida F. Schmidt var. anhuiensis (Y. K. Yang, N. Wang & W. C. Ye) Ziman & Dutton in Fl. China 6: 311. 2001; Ziman et al. in J. Jap. Bot. 79 (5): 298. 2004. Type: 安徽,青阳,九华山,天台,alt. 1 000 m,山谷松林下,1987-04-16,叶文才87103 (holotype, WUBI; isotype, PE), 87101, 87102,  87114, 87125, 87126,87127 (paratype, WUBI)。
  特產于中国安徽省青阳县九华山。生于山谷松林下,海拔1 000 m。
  系4.  加查银莲花系
  Ser.  Jiachaenses W. T. Wang, ser. nov. Type: A. jiachaensis W. T. Wang.
  A sectionis hujus seriebus ceteris differt foliis basalibus duodecium per plantam. In seriebus ceteris, folia basalia unum ad tria, raro quatuor ad septem per plantam sunt.
  Rhizome not putting forth a stolon. Basal leaves ca. 12, 3-sect. Cymes 1-3-flowered. Sepals 5. Anthers on apex obtuse. Carpels ca. 70, glabrous; styles absent; stigmas globose.   本系的每一株具约12枚基生叶,而与本组其他系不同。
  根状茎不生出匍匐茎。基生叶约12,3全裂。聚伞花序有1~3花。萼片5。花药顶端钝。心皮约70,无毛;花柱不存在;柱头球形。
  1种。
  69.  加查银莲花 图版25: 8-11
  Anemone jiachaensis  W. T. Wang in Pl. Divers. Resour. 36(4): 450, fig. 1: H-K. 2014. Holotype 西藏:加查,近县城,alt. 3 400 m,山谷阶地,花白色,1980-08-02,倪志诚,次多,次旦2734 (PE)。
  特产于中国西藏加查。生于山谷阶地上,海拔3 400 m。
  組7.  直果银莲花组
  Sect. Pleurocarpus W. T. Wang, sect. nov Type: A. orthocarpa Hand. -Mazz.
  Ob achenia glabra 1-vel 4-costata sectio nova haec est similis Sect. Begoniifoliae Tamura, a qua carpellorum stylis brevioribus vel longioribus conspicuis, stigmatibus inconspicuis differt.
  Rhizomes subvertical, short and thick. Leaf blades undivided, 3-lobulate or 3-parted. Cymes 1-flowered. Sepals 5, white. Stamen filaments filiform; pollen grains 6-colpate or pantoporate. Carpels glabrous; styles or short, or longer, conspicuous; stigmas inconspicuous. Achenes longitudinally 1- or 4-ribbed. Chromosome basic number x=8.
  此组的瘦果无毛,具1条或4条肋,而与卵叶银莲花组(Sect. Begoniifolia)相似,但本组的心皮具明显的短或较长花柱,柱头不明显,而与卵叶银莲花组不同。
  根状茎近垂直,短,粗。叶片不分裂,3微裂或3深裂。聚伞花序具1花。萼片5,白色。雄蕊花丝丝形;花粉具6沟或具散孔。心皮无毛;花柱短或较长,明显;柱头不明显。瘦果具1条或4条纵肋。染色体基数=8(Zhang et al., 2015)。
  2种。
  系1.  兴义银莲花系
  Ser. Xingyienses W. T. Wang, ser. nov. Type: A. xingyiensis Q. Yuan & Q. E. Yang.
  Folia basalia 9-20; laminae ellipticae, basi cuneatae, indivisae vel inconspicue 3-lobulatae. Pollinis grana pantoporata. (Zhang et al., 2015) Carpellorum styli breves. Achenia longitudinaliter 1-costata.
  1种。
  70. 兴义银莲花 图版26
  Anemone xingyiensis Q. Yuan & Q. E. Yang in Bot. Stud. Taipei 50: 493, figs. 1-2. 2009; Q. E. Yang in Plant Syst. Evol. 301: 421 et 422. 2015. Type 贵州:兴义,Malinghe,山谷,alt. 1 000 m,喀斯特陡崖湿处,2007-03-25,袁琼,杨亲二1077 (holotype, IBSC, non vidi);同地,2007-04-21,袁琼,杨亲二,刘英678 (paratype, IBSC, non vidi)。
  特产于中国贵州省兴义市。生于石灰岩山峭壁上湿处,海拔1 000 m。
  系2. 直果银莲花系
  Ser. Orthocarpae W. T. Wang, ser. nov. Type: A. orthocarpa Hand. -Mazz.
  Folia basalia 4-6; laminae pentagonae, basi vadose cordatae, 3-partitae. Pollinis grana 6-colpata. (Zhang et al., 2015). Carpellorum styli longiores, subulati, recti vel apice curvati. Achenia longitudinaliter 4-costatae.
  1种。
  71. 直果银莲花 图8
  Anemone orthocarpa Hand. -Mazz. in Acta Hort. Gotob. 13: 176. 1939; W. T. Wang in Fl. Reip. Pop. Sin. 28: 26. 1980; Y. K. Li in Fl. Guizhou. 3: 80. 1990; Wang, Ziman & Dutton in Fl. China 6: 319. 2001; Ziman et al. in J. Jap. Bot. 83(3): 149. 2008; Q. E. Yang in Plant Syst. Evol. 301: 422. 2015. Syntypes 贵州西南部:兴义,Hoang-tsao-pa, Tien-hsin-chiao (黄草坝,天仙桥),1915-02-27,J. P. Carvalerie 4599 (P, K; isosyntype, PE)。   分布于中国贵州省兴义市以及越南(Zhang & Yang, 2014)。
  组8. 卵叶银莲花组
  Sect. Begoniifolia (Ulbr.) Tamura in Sci. Rep. Osaka Univ. 16: 26. 1967; W. T. Wang in Fl. Reip. Pop. Sin. 28. 25. 1980; Tamura in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17a IV: 335. 1995; Ziman et al. in J. Jap. Bot. 83(3): 18, 148. 2008. — Sect. Pulsatilloides ser. Begoniifolia Ulbr. in Bot. Jahrb. 37: 202, 243. 1906. — Pulsatilloides sect. Begoniifolia (Ulbr.) Starod. Syst. & Evo. 124. 1991. —Sect. Rivularium subsect. Begoniifoliae (Ulbr.) Q. E. Yang in Plant Syst. Evol. 301: 421. 2015, p. p. excl. A. orthocarpa Hand. -Mazz. et A. xingyiensi Q. Yuan & Q. E. Yang. Type: A. begoniifolia Lévl. & Van.
  根狀茎近垂直。基生叶叶片不分裂或3~5浅裂,稀3全裂。聚伞花序伞状,有1~7花;总苞苞片无柄或有柄,不分裂或3全裂。萼片5,白色,粉红或紫色。雄蕊花丝丝形;花粉具3或6沟,或具散孔(Zhang et al., 2015)。心皮无毛;花柱短,不明显;柱头明显。瘦果无毛,具4条细纵肋。染色体基数x=8(Hoot et al., 1994; Zhang et al., 2015)。
  约6种。分布于中国西南部,以及缅甸、马来西亚、印度尼西亚的苏门答腊岛。中国有4种,分布于西南山区。
  72.  卵叶银莲花 图版15: 1-5
  Anemone begoniifolia Lévl. & Van. in Bull. Acad. Geogr. Bot. 9: 46. 1902; Finet & Gagnep. in Bull. Soc. Bot. France 51: 71. 1904; Ulbr. in Bot. Jahrb. 37: 243. 1905; Pei in Contr. Biol. Lab. Sci. Soc. China, Bot. ser., 9(1): 5. 1933; Hand. -Mazz. in Acta Hort. Gotob. 13: 179. 1939; Iconogr. Corm. Sin. 1: 727, fig. 1454. 1972; W. T. Wang in Fl. Reip. Pop. Sin. 25: 25, pl. 12: 1-5. 1980; Y. K. Li in Fl. Guizhou. 3: 78. 1990; W. T. Wang
  in Fl. Guangxi 1: 278, pl. 119. 5. 1991; in Fl. Yunnan. 11: 190, pl. 57: 1-5. 2000; et in High. Pl. China 3: 493, fig. 786. 2000; Wang, Ziman & Dutton in Fl. China 6: 319. 2001; Ziman et al. in J. Jap. Bot. 83(3): 150. 2008; Q. E. Yang in Plant Syst. Evol. 301: 421. 2015; L. Xie in Medic. Fl. China 321. 2016. Holotype贵州:Touchan 独山,1899-04,J. P. Cavalerie 2600 (non vidi)。
  A. esquirolii Lévl. in Repert. Sp. Nov. 9: 446. 1911.
  分布于中国云南东南部(海拔1 000~1 800 m)、广西、贵州(550~1 260 m)、四川东南、重庆南部(300~780 m)。生于山地草坡、陡崖上或林中,海拔300~1 800 m。
  本种的根可供药用,有消肿接骨、止血生肌之效,用于风湿关节痛、疮毒等症(谢磊等,2016)。
  标本登录 云南:富宁,王启无87126,88707;文山,蔡希陶51762;麻栗坡,王启无86254,税玉民,陈文红20311,金效华,刘冰等1017。广西:?297。贵州:安龙,张志松,张永田2523;绥阳,钟补勤417;德江,罗毅波502。四川:筠连,傅德志84-331。重庆:南川,曲桂岭1685,熊济华,周子林90099,李国凤60322,傅德志84-03;武隆,刘正宇182536。
  73. 河口银莲花 图版29: 8
  Anemone hokouensis C. Y. Wu ex W. T. Wang in Acta Phytotax. Sin. 12(2): 168, pl. 9: 9. 1974; in Fl. Reip. Pop. Sin. 28: 26, pl. 7: 9: 1980; et in Fl. Yunnan. 11: 190. 2000; Wang, Ziman & Dutton in Fl. China 6: 319. 2001; Ziman et al. in J. Jap. Bot. 83(3): 150. 2008; Q. E. Yang in Plant Syst. Evol. 301: 421. 2015. Holotype 云南河口,薄竹箐,alt. 1 200 m, 1953-05-31,阙再旦1237 (KUN)。   特产于中国云南河口。生于海拔1 200 m一带山地。
  74. 拟卵叶银莲花 图版29: 4-7
  Anemone howellii J. F. Jeff. & W. W. Smith in Not. Bot. Gard.Edinb. 8: 78. 1916; W. T. Wang in Fl. Reip. Pop. Sin. 28: 25, pl. 7: 4-7. 1980; in Bull. Bot. Res. Haibin 3(1): 36. 1983; in Fl. Guangxi 1: 278. 1991; et in Fl. Yunnan. 11: 189. 2000; Wang, Ziman & Dutton in Fl. China 6: 319. 2001; Ziman et al. in J. Jap. Bot. 83(3): 150. 2008; Q. E. Yang in Plant Syst. Evol. 301: 422. 2015. Holotype 云南:腾冲,E. B. Howell 111 (E; photo, PE)。
  A. begoniifolioides W. T. Wang in Acta Phytotax. Sin. 12(2): 167. 1974. Type 云南:文山,alt. 2 300 m, 1947-08-17,冯国楣17299 (holotype, PE);马关,蔡希陶57874 (paratype, PE);屏边,王孝100238 (paratype, IBSC);河口,胡月英143 (paratype, YIM)。
  分布于中国云南西部和东南部、贵州西南部以及缅甸北部。生山谷沟边或林中,海拔1 200~2 300 m。
  标本登录 云南:麻栗坡,金效华,刘冰等1017,王印政等4177,税玉民,方瑞正,陈文红21218;屏边,瑶山区,毛品一4287。贵州:安龙,张志松,张永田2913。
  75.  盘蕊银莲花 图版27
  Anemone discoideogyna W. T. Wang, sp. nov. Type 四川 (Sichuan):石棉县,金窝 (Shimian Xian, Jinwo), alt. 2 179 m,灌丛中,花白色 (in bushes, fls. white), 2018-07-18,朱相云,张若鹏,祖奎玲 (X. Y. Zhu, R. P. Zhang & K. L. Zu) G10222 (holotype and isotype, PE)。
  Herbs perennial, 34-45 cm tall. Basal leaves 2-3, 1 simple, the other 1 or 2 compound, 2-3-leafletted, all long petiolate; simple leaf with blade cordate, 6-11 × 5.4-12 cm, 3-lobed, on margin denticulate, adaxially appressed-puberulous, abaxially on nerves with sparse short hairs, and with petiole 7.4-10.5 cm long; compound leaves ternate or 2-leafletted, with lateral leaflets smaller than terminal one, broadly and obliquely ovate, and with petioles 5.5-8 cm long. Scapes 2, 15-21 cm long, with petioles densely puberulous; cymes terminal, 2-7-flowered; involucral bracts 2, shortly petiolate, deltoid-ovate, 2-3 × 2-3 cm, often 3-lobed, on margin denticulate; pedicels 4-14 cm long, densely appressed-puberulous. Besides, a single flower with its pedicel ca. 24 cm long, directly arising from the rhizome apex. Flower: Sepals 5, white, broadly obovate or obovate, 1.9-2.5 × 0.9-1.9 cm, at apex rounded, adaxially glabrous, abaxially densely puberulous. Stamens ca. 240, glabrous; filaments filiform, 2-7 mm long; anthers oblong, 1.5-2 × 0.5-0.8 mm. Gynoecium disc-like, 6 mm in diam., consisting of ca. 200 compactly aggregate carpels; carpel 1.5 mm long, with puberulous ovary broad-elliptic, ca. 0.5 mm long and sessile subglobose stigma ca. 0.4 mm in diam., and with a glabrous stipe ca. 0.6 mm long.
  特產于中国四川石棉。生于山地灌丛中,海拔2 179 m。   76. 墨脱银莲花 图版25: 1-4
  Anemone motuoenis W. T. Wang in Pl. Divers. Resour. 36(4): 449, fig. 1: A-D. 2014. Holotype 西藏:墨脱,那格之西,alt.3 600 m,冷杉林中,花粉红色,1980-06-25,青藏高原组10307 (PE)。
  特产于中国西藏墨脱县。生于山地冷杉林中,海拔3 600 m。
  组9.  二岐银莲花组
  Sect.Anemonidium Spach, Hist. Nat. Veget. Phan. 7. 1839; Jancz. in Rev. Gen. Bot. 4: 253. 1892; Ulbr. in Bot. Jahrb. 37: 206, 261. 1905-06; Tamura in Sci. Rep. Osaka Univ. 16(2): 27. 1967; W. T. Wang in Fl. Reip. Pop. Sin. 28: 56. 1980; Tamura in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17AIV: 333. 1995; Ziman et al. in J. Jap. Bot. 83(3): 139, 150. 2008. Type: A. dichotoma L. — Anemonidium (Spach) Holub in Folia Geobot, Phytotax. Praha 9: 272. 1974. — Anemonastrum Holub sect. Anemonidium (Spach) Mosyakin in Phytoneuron 79: 4. 2016.
  根状茎细长。基生叶1~2或不存在,具长柄。聚伞花序2~3回二歧状分枝;总苞苞片2或3,无柄,小总苞苞片2,无柄。萼片5或6。雄蕊花丝丝形;花粉具3沟。瘦果扁平,每侧具翅。染色体基数x=7(Ehrendorfer et al., 2001)。
  2种,分布于亚洲、欧洲和北美洲温带地区。中国有1种,分布于吉林、黑龙江和内蒙古。
  77. 二岐银莲花 图版28: 1-4
  Anemone dichotoma L. Sp. Pl. 540. 1753; DC. Prodr. 1: 20. 1824; Finet & Gagnep. in Bull. Soc. Bot. France 51: 73. 1904; Ulbr. in Bot. Jahrb. 37: 261. 1906; Juz. in Fl. URSS 7: 268. 1937; Iconogr. Corm, Sin. 1: 731, fig. 1462. 1972; Fl. Pl. Herb. China Bor. -Or. 3: 156. 1975; W. T. Wang in Fl. Reip. Pop. Sin. 28: 56, pl. 16: 1-4. 1980; Y. Z. Zhao in Fl. Intramongol., 2nd ed., 2: 463, pl. 187: 1-2. 1991; W. T. Wang in High. Pl. China 3: 499, fig. 799. 2000; Wang, Ziman & Dutton in Fl. China 6: 310. 2001; Ziman et al. in J. Jap. Bot. 83(3): 150. 2008; Y. Z. Zhao, Class. Distr. Vasc.  Pl. Inn. Mongol. 162. 2012. — Anemonastrum dichotomum (L.) Holub in Folia Geobot. Phytotax. Praha 9: 272.1974. — Anemonastrum dichotomum (L.) Mosyakin in Phytoneuron 79: 4, 2016.  Described from Siberia.
  分布于中國吉林、黑龙江和内蒙古,以及亚洲北部和欧洲。生于丘陵或低山山坡湿草地,河边,沼泽中或林中,海拔450~980 m。
  本种的根状茎可供药用,有舒筋活血、清热解毒、止痢之效,用于跌打损伤,风湿性关节炎、痢疾等症(谢磊等,2016)。
  标本登录 吉林:和龙,延边二组563;敦化,孔宪武1620。黑龙江:镜泊湖,安部坚太9793;尚志,P. Licent 8879;哈尔滨,王薇等518;呼兰,P. Licent 8898;北安,东北队267;伊春,中德队7708;富锦,二龙山,张玉良1635。内蒙古:乌兰浩特,叶建飞09-62811;呼伦贝尔,王战等641,傅沛云1898;牙克石,东北林学院队86-23;大兴安岭,1954-08,林型组s. n.
  组10. 打破碗花花组
  Sect.Anemone. W. T. Wang in Fl. Reip. Pop. Sin. 28: 27. 1980. —Subgen. Anemone. Tamura in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17aIV: 344. 1995.
  基生叶少数或数枚,具长柄。萼片5~10。花药宽椭圆形或长圆形;花丝丝形,或下部稍宽;花粉具3沟具散孔或具散沟,稀具螺旋状萌发孔(Xi & Chang, 1964; Ehrendorfer et al., 2009)。心皮数目常极多。瘦果小,数目也常极多,形成球形体、卵球形体或圆筒状体,密被长绵毛,无柄或具柄。染色体基数x=8(Tamura 1995;Ehrendorfer et al., 2001)。   约40种,分布于亚洲、欧洲和非洲北部,以及北美洲和南美洲。中国有8种,广布于西部、中部和东北。
  系1.  岩生银莲花系
  Ser. Rupicolae Tamura ex Chaudhary & Trifonova in Bot. Zhurn. 73: 808. 1988; Tamura in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17a IV: 346. 1995. — A. subsect. Rupicolae (Tamura ex Chaudhary & Trifonova) Starod., Anemone: Syst. Evol. 119. 1991; Q. E. Yang in Plant Syst. Evol. 301: 421. 2015. Type: A. rupicola Camb.
  A.sect. Eriocephalus Hook. f. & Thoms. Fl. Ind. 20. 1855; Tamura in I. c. 345, p. p. Leptotype: A. rupicola Camb.
  根状茎不成块状。聚伞花序有1~7花,简单,不分枝,或分枝;总苞苞片通常3,与基生叶相似。萼片5~10。瘦果具短柄或无柄。
  约16种。分布于亚洲、欧洲、北美洲和南美洲。中国有4种。
  77. 大花银莲花 图版30: 7-9
  Anemone sylvestris L. Sp. Pl. 540. 1753;DC. Prodr. 1: 20. 1824; Finet & Gagnep. in Bull. Soc. Bot. France 51: 89. 1904; Ulbr. in Bot. Jahrb. 37: 254. 1905; Juz. in Fl. URSS 7: 266. 1937; Iconogr. Corm. Sin. 1: 729, fig. 1957. 1972; W. T. Wang in Fl. Reip. Pop. Sin. 28: 33, pl. 6: 7-9. 1980; S. Y. He, Fl. Beijing, rev. ed., 1: 617, fig. 392. 1984; J. W. Wang in Fl. Hebei. 1: 464, fig. 466. 1986; Y. Z. Zhao in Fl. Intramongol., 2nd ed., 2: 685, pl. 187: 3-5. 1991; X. Y. Yu, B. L. Li & Y. M. Li in Fl. Shanxi. 1: 617, pl. 392. 1992; J. G. Liu in Fl. Xinjiang 2(1): 277, pl. 75: 4-6. 1994; W. T. Wang in High. Pl. China 3: 495, fig. 490. 2000; Wang, Ziman & Dutton in Fl. China 6: 317. 2001; Ziman et al. in J. Jap. Bot. 83(3): 142. 2008; Y. Zhou, Colour. Ill. Pl. Wet Area. NE China 334. 2019. Described from Germany.
  分布于中國河北北部、辽宁西部、吉林西部、黑龙江、内蒙古和新疆,以及亚洲北部和欧洲。生于草原、山地草坡、林边或多沙坡地,海拔580~2 000 m。
  标本登录 河北:张家口,1972-06,乌兴曾s. n.;围场,傅国勋36。北京:延庆,1971-06,延庆队158。内蒙古:大青山,马毓泉,吴庆如131;克什克腾旗,杨雅玲277,陈又生14-656;索伦,刘慎谔8249;阿尔山,张玉良350;博客图,J. Sato 90;呼伦贝尔,王战等543,770。新疆:哈密,林型组队16;富蕴,新疆队10387;博格达山,新疆学院植物队208;乌鲁木齐,西北植物所新疆队147;天山,天池,王忠涛29;阜康,洪德元等186;沙湾,关克俭1278,2302;托里,许小纯93-127;绥定,周太炎等65-856;霍城,李安仁,朱家柟10418。
  79. 岩生银莲花 图版2: 6-8
  Anemone rupicola Camb. in Jacq. Voy. 4: 5, fig. 2. 1838; Hook. f. & Thoms. in Hook. f. Fl. Brit. Ind. 1: 8, 1872; Franch in Bull. Soc. Bot. France 33: 364. 1886; et Pl. Delav. 7: 1889; Finet & Gagnep. in Bull. Soc. Bot. France 51: 65. 1904; Ulbr. in Bot. Jahrb. 37: 256. 1905; Pei in Contr. Biol. Lab. Sci. Soc. China, Bot. Ser., 9(1): 5, 1933; Hand. -Mazz. in Acta Hort. Gotob 13: 179. 1939; W. T. Wang in Fl. Reip. Pop. Sin. 28: 11, pl. 5: 6-8. 1980; in Fl. Xizang. 2: 80, fig. 22: 3. 1985; in Fl. Yunnan. 11: 193. 2000; et in High. Pl. China 3: 494, fig. 789. 2000; Wang, Ziman & Dutton in Fl. China 6: 318. 2001; Ziman et al. in J. Jap. Bot. 83(3): 142. 2008. Described from the Himalayan region.   分布于中国云南西北部、四川西部、西藏东部和南部,以及不丹、尼泊尔和印度北部。生于山地多石砾山坡、草地、灌丛或林下,海拔2 400~4 600 m。
  标本登录 云南:德钦,alt. 2 800~4 500 m,王启无69463,俞德浚8441,8743,9069,9354,青藏队2750;中甸,俞德浚11574,12323,冯国楣1254;丽江,俞德浚8724,15207,秦仁昌31002。四川:木里,俞德浚5831,6447,青藏队14306;乡城,周仕顺6065;得荣,D. E. Boufford等30830;冕宁,武素功2268;康定,刘振书1504;Minya Konka雪山,alt. 4 600 m,1909-06,J. F. Rock 17677;宝兴,俞德浚2285,曲桂龄3572,张秀实,任有铣5483。西藏:察隅,青藏队271,1085;波密,应俊生,洪德元5853;聂拉木,张永田,郎楷永3885,3925,4252;吉隆,吴征镒等75-674,西藏中草药队533,685。
  80. 锐裂银莲花 图9
  Anemone laceratoincisa W. T. Wang in Fl. Reip. Pop. Sin. 28: 33, 349, fig. l. 1980; Wang, Ziman & Dutton in Fl. China 6: 318. 2001; Ziman et al. in J. Jap. Bot. 83(3): 142. 2008. —A. rupicola Camb. ssp. laceratoincisa (W. T. Wang) Chaudhary in Bot. Zhurn. 72(6): 824. 1987, et 73(8): 1200. 1988. Holotype  甘肃:文县,燕子牙,?125 (CMMI, photo, PE).
  特产于中国甘肃文县。
  81. 西藏银莲花 图版29: 1-3
  Anemone tibetica W. T. Wang in Fl. Reip. Pop. Sin. 28: 32, 349, pl. 7: 1-3. 1980; et in Fl. Xizang. 2: 80, fig. 23: 1-2. 1985; Wang, Ziman & Dutton in Fl. China 6: 38. 2001; Ziman et al. in J. Jap. Bot. 83(3): 142. 2008; W. T. Wang in High. Pl. China in Colour 3: 404. 2016. Holotype 西藏:朗縣,泵村,alt.3 100 m,沟边,1972-08-18,西藏中草药队4423 (PE)。
  特产于中国西藏朗县。生于山谷沟边,海拔3 100 m。
  系2. 打破碗花花系
  Ser. Anemospermos DC. ex Ulbr. in Bot. Jahrb. 37: 205. 1905; Chaudhary & Trifonova in Bot. Zhurn. 73: 808. 1988. Lectolype: A. vitifolia Buch.-Ham. ex DC.
  Eriocapitella Nakai in J. Jap. Bot. 17: 267. 1941. — A. sect. Eriocapitella (Nakai) Tamura in Acta Phytotax. Geobot. 42: 180, 1991; et in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17aIV: 344. 1995. Type: E. japonica (Thumb.) Nakai.
  A. subsect. Vitifoliae Starod. in Bot. Zhurn. 74: 1345. 1989; Q. E. Yang in Plant Syst. Evol. 301: 421. 2015. Type: A. vitifolia Buch. -Ham. ex DC.
  根状茎垂直,不成块状。基生叶为三出复叶或为单叶。聚伞花序2~4回分枝;总苞苞片3,具柄,为三出复叶或单叶。萼片5。雄蕊花丝丝形。心皮400~500。瘦果小,密被长绵毛,有细长柄。
  3种。分布于中国西南、长江流域和北部诸省,以及尼泊尔、缅甸和印度北部。
  82. 打破碗花花 图版30: 1-3
  Anemone hupehensis (Lem.) Lem. Cat. 176: 40. 1910; Bowles & Stearn in J. Roy.Hort. Soc. 72: 261. 1947; Icongr. Corm. Sin. 1: 728, fig. 1455. 1972; Fl. Tsinling. 1(2): 282. 1974; W. T. Wang in Fl. Reip. Pop. Sin. 28: 27, pl. 6: 1-3. 1980; B. Z. Ding et al. Fl. Henan. 1: 476. 1981; X. W. Wang in Fl. Anhui. 2: 327, fig. 631. 1987; Y. K. Li in Fl. Guizhouo. 3: 76, pl. 3: 1-6. 1990; W. T. Wang in Fl. Guangxi 1: 280, pl. 119. 5. 1991; Z. H. Lin in Fl. Zhejiang 2: 280, fig. 2-371. 1992; K. M. Liu, Fl. Hunan. 2: 660, fig. 2-516. 2000; W. T. Wang in High. Pl. China 3: 493, fig. 787. 2000; S. H. Fu, Fl. Hubei. 1: 350, fig. 487. 2001; Wang, Ziman & Dutton in Fl. China 6: 317. 2001; W. T. Wang in Fl. Jiangxi 2: 160, fig. 160. 2004; Ziman et al. in J. Jap. Bot. 83(3): 141. 2008; L. Xie in Medic. Fl. China 3: 322. 2016. —A. japonica (Thunb.) Sieb. & Zucc. var. hupehensis Lem. Cat. 170: 42. 1908. Described from Hubei Province.   82a. var. hupehensis
  82ai. f. hupehensis
  分布于中国四川、云南、广西北部、广东北部、贵州、重庆、甘肃南部、陕西南部、河南南部、湖北、湖南、江西、安徽南部和浙江。生于丘陵或低山草坡或沟边,海拔400~1 800 m。
  本种的根状茎和全草可供药用,有消肿利湿、消肿散瘀之效,用于消化不良、肠炎、痢疾、疟疾、瘰疬等症; 有小毒(谢磊等,2016)。全草可用作土农药,水浸液可防治稻苞虫、稻螟、棉蚜、蝇蛆等。
  标本登录 四川:木里,青藏队13308;普格,植物资源队59-5542;雷波,赵清盛1483;美姑,植物资源队59-1364;马边,俞德浚5861;峨眉山,?2561;都江堰,朱大海952;汶川,郎楷永,李良千,费勇1372;茂县,汤宗孝1507;平武,方文培4438;苍溪,经济植物队59-5030;万源,李培元4426。云南:维西,杨亲二94-70;丽江,横断山队2685;文山,税玉民336;西畴,冯国楣11983。广西:那坡,王洪6588;凌云,秦仁昌6838;天娥,北京队83-3196;九万山,陈少卿15911;兴安,李光照15325;龙胜,广福林区队1076。广东:地址不明,1928,蒋英1327。贵州:兴义,安顺队650;安龙,张志松,张永田3622;册亨,曹子余532;纳雍,毕节队304;毕节,禹平华444;金沙,黄昌洪58;习水,毕节队1550;赤水,毕节队1352;遵义,川黔队1175;清镇,川黔队1694;贵阳,安明态5288;独山,荔波队1444;凯里,黔南队3450;石阡,武陵山队2994;梵净山,简焯坡等30355。重庆:缙云山,缙云山队171;南川,金佛山,熊济华,周子林92741;武隆,刘正宇181309;酉阳,赵佐成2853;开县,巴山队2551;奉节,周洪富,粟和毅110866;巫山,周洪富109857;巫溪,杨光辉59265;城口,戴天伦105896。甘肃:文县,何业祺911;康县,张志英16743。陕西:略阳,傅坤俊6042;勉县,傅坤俊3890;洋县,郭本兆1844;紫阳,李培元4911;平利,刘继孟8476;周至,刘继孟8740。河南:淅川,林业厅队1460。湖北:来凤,李洪钧7197;鹤峰,李洪钧6356;利川,傅国勋1572;巴东,陈权龙878;秭归,赵常明931;兴山,王作宾11967;神农架,神农架队11405。湖南:新宁,刘林翰152111;武冈,云山,刘林翰20069;芷江,武陵山队2013;永顺,刘林翰9508;桑植,北京队4358;石门,壶瓶山队A214;东安,刘瑛826。江西:莲花,杨祥学1344;武功山,江西队54-992;修水,熊济水99-12。安徽:绩溪,叶文才88-101。浙江:天台山,贺贤育28254;开化,丁炳杨1250。
  82aii. 水棉花
  f. alba W. T. Wang in Acta Phytotax. Sin. 12(2): 166. 1974; in Fl. Reip. Pop. Sin. 28: 228. 1980; et in Fl. Yunnan. 11: 191. 2000; Wang, Ziman & Dutton in Fl. China 6: 317. 2001; W. T. Wang in High. Pl. China in Colour 3: 402. 2016. Holotype 四川:宝兴,盐井,alt.1 300 m, 1963-09,关克俭,王文采13111 (PE)
  Paratypes (均存PE) 四川:布拖,胡嘉琪等5529;会理,俞德浚1366。云南:昆明,毛品一77;大理,蔡希陶58390;德钦,俞德浚10577;镇雄,禹平华978。
  A. vitifolia Buch. -Ham. ex DC. var. matsudai Yamamoto, Suppl. Tc. Pl. Formos. 3: 27. 1906. — A. matsudai (Yamamoto) Tamura in Acta Phototax Geobot. 16(4): 110. 1956.
  A. hupehensis var. simplicifolia W. T. Wang in Acta Phytotax. Sin. 12(2): 167. 1974. Type 四川:寶兴,硗碛,alt. 2 400 m,1963-09-06,关克俭,王文采3071 (holotype, PE);峨眉山,九老洞,1952-09-01,熊济华等32288 (paratype, PE)。
  ?A. elegans Decne. in Rev. Hort. 4 ser., 1: 41, tab. 3. 1852; Hand.-Mazz. in Acta Hort. Gotob. 13: 179. 1939. — A. japonica Sieb. & Zucc. var. elegans (Decne.) Franch. in Bull. Soc. Bot. France. 33: 364. 1886; et Pl. Delav. 9. 1890; Ulbr. in Bot. Jahrb. 37: 253. 1905.
  A. vitifolia auct. non Buch. -Ham. ex DC.: T. Y. Yang & T. C. Huang in Fl. Taiwan, 2nd ed., 2: 509, pl. 236. 1996.
  分布于中国云南(海拔1 400~3 500 m)、四川西部(900~2 400 m)、贵州西部(1 500~1 980 m)、重庆南部和台湾(1 400~2 300 m)。生于山地草坡,灌丛,沟边,林边或林下。   标本登录 云南:贡山,俞德浚22693;德钦,王启无69127;中甸,俞德浚10942,冯国楣2053;维西,蔡希陶57863,王启无58292;福贡,蔡希陶57079;丽江,秦仁昌21261;鹤庆,秦仁昌23367;漾濞,金效华1339;大理,王汉臣1415;禄劝,毛品一1562;富民,邱炳云596018;昆明,钟观光2128,刘慎谔14893;屏边,王启无82117;东川,蓝顺彬253;大关,滇东北队484。四川:峨眉山,杜大华648;石棉,赵佐成3273;泸定,D. E. Boufford等27409;天全,胡文光,何铸11857。贵州:兴义,安顺队650;盘县,安顺队1222;关岭,王文采s. n.; 纳雍,毕节队446;大方,毕节队921;黔西,蒋英8827。重庆:南川,金佛山,熊济华,周子林93745。台湾:南投,C. M. Wang et al. 3309;花莲,D. E. Boufford等19285,杨宗愈2183;桃园,杨宗愈6251。
  82b. 秋牡丹 (变种)
  var. japonica (Thunb.) Bowles & Stearn in J. Roy. Hort. Soc. 72: 265. 1947; W. T. Wang in Fl. Reip. Pop. Sin. 28: 28. 1980; X. R. Lin & X. Z. Zhao in Fl. Fujian. 2: 21. 1985; X. W. Wang in Fl. Anhui 2: 327. 1987; W. T. Wang in Fl.Guangxi 1: 280. 1991; Z. H. Lin in Fl. Zhejiang 2: 280, fig. 2-372. 1992; K. M. Liu, Fl. Hunan 2: 660. 2000; Wang, Ziman & Dutton in Fl. China 6: 317. 2001; R. J. Wang in Fl. Guangdong 5: 17. 2004; W. T. Wang in Fl. Jiangxi 2: 160. 2004; M. B. Deng & K. Ye in Fl. Jiangsu 2: 87, fig. 2: 141. 2013. — Atragene japonica Thunb. Fl. Jap. 239. 1784. — Anemone japonica (Thunb.) Sieb. & Zucc. Fl. Jap. 1: 15. 1885, non Houtt, 1778; Forbes & Hemsl. in J. Linn. Soc. Bot. 23: 11. 1886; Ulbr. in Bot. Jahrb. 37: 252. 1905; Chun in Sunyatsenia 1: 228. 1934, p. p. — A. vitifolia var. japonica (Thunb.) Finet & Gagnep. in Bull. Soc. Bot. France 51: 68. 1904. — Eriocapitella japonica (Thunb.) Nakai in J. Jap. Bot. 17: 268. 1941. Described from Japan.
  A. scabiosa Lévl. & Van.in Bull. Acad. Geogr. Bot. 11: 47. 1902.
  为一栽培品种,在中国长江流域及以南诸省区广泛栽培,有时逸出呈野生状。在日本也有。
  83. 大火草
  Anemone tomentosa (Maxim.) Pei in Contr. Biol. Lab. Sci. Soc. China, Bot. Ser. 9: 2. 1933; et Icon. Chin. Medic. Pl. 1: fig. 17. 1955; X. W. Kung, Fl. Lanzhou 252, fig. 187. 1962; Iconogr. Corm. Sin. 1: 728, fig. 1456. 1972; Fl. Tsinling 1(2): 283, fig. 243. 1974; J. W. Wang in Fl. Hebei 1: 464, fig. 467. 1986; D. Z. Ma & H. L. Liu, Fl. Ningxia 1: 191. 1986; X. Y. Yu, B. L. Li & Y. M. Li in Fl. Shanxi 1: 617, fig. 391. 1992; L. H. Zhou in Fl. Qinghai. 1: 339. 1997; S. H. Fu, Fl. Hubei. 1: 350, fig. 488. 2001; Wang, Ziman & Dutton in Fl. China 6: 317. 2001; Ziman et al. in J. Jap. Bot. 83(3): 141. 2008; L. Xie in Medic. Fl. China 3: 324. 2016. —A. japonica var. tomentosa Maxim. Fl. Tangut. 7. 1889; et in Acta Hort. Petrop. 11: 20. 1890; Ulbr. in Bot. Jahrb. 37: 253. 1905. — A. vitifolia var. tomentosa (Maxim.) Finet & Ggagnep in Bull. Soc. Bot. France 51: 69. 1904. — A. elegans Decne. var. tomentosa (Maxim.) Hand.-Mazz. in Acta Hort. Gotob. 13: 179. 1939. — Eriocapitella vitifolia var. tomentosa (Maxim.) Nakai in J. Jap. Bot. 17: 270. 1941. Syntypes: 甘肅,1875,P. J. Piasetski s. n. (non vidi);同地,1885, G. N. Potanin s. n. (LE, non vidi, photo, PE)。   分布于中国四川西部(康定以北,海拔2 000~3 400 m)、重庆(1 700~1 750 m)、湖北西部(500~2 200 m)、青海东部(1 900~2 500 m)、甘肃南部(1 350~1 800 m)、陜西(780~1 800 m)、河南西部(360~2 000 m)、山西(1 100~1 800 m)和河北西部(700~800 m)。生于山地草坡,灌丛,河滩,林边或林下。
  本种的根可供药用,有化痰、散瘀、消食化积之效,用于劳伤咳喘、跌打损伤、小儿疳积、痢疾等症; 有小毒(谢磊等,2016)。
  标本登录 四川:雅江,姜恕2673;康定,方文培3637;二郎山,关克俭1875;宝兴,张秀实,任有铣6772;小金,郎楷永,李良千,费勇1526;金川,李馨76135;丹八,D. E. Boufford等37947;马尔康,李馨71929;理县,赵清盛252;平武,鲁丽敏2008-365;松潘,傅坤俊1768;九寨沟,李沛琼57;若尔盖,朱大海4411。重庆:开县,戴天伦101753;巫溪,曲桂龄1738;城口,戴天伦101285。湖北:巴东,傅国勋,张志松891;神农架,236-6队2905。青海:循化,周立华s. n.;民和,何廷农825;互助,郭本兆9505。甘肃:文县,郝景盛374;康县,廉永善96-398;舟曲,何业祺688;临潭,王作宾6138;兰州,何业祺4945;武山,黄河队4615;天水,刘继孟10450;华亭,王作宾16939;平凉,陕甘队10110。陕西:洋县,杨金祥769;宁陕,傅坤俊17622;佛坪,傅坤俊4950;太白山,王作宾1503;终南山,白银元1171;蓝田,苏贵兴122;华山,郝景盛4176;黄龙山,傅坤俊3092;延安,傅坤俊8149;志丹县,崔友文10821;靖边,黄河队8035。河南:固始,刘继孟4723;内乡,关克俭882;西峡,关克俭1007;栾川,马成功367;卢氏,河南队34483;嵩县,植物资源队15;登封,普查队54501;林县,普查队4025。山西:芮城,包世英,严生俊1548;夏县,刘天慰447,陵川,刘继孟7630;霍州,黄河队851;灵石,刘鑫源21136;中阳,黄河队2090;交城,刘鑫源21358;兴县。黄河队2638;芦芽山,Harry Smith 8157;五台,关克俭,陈艺林1989;繁峙,段长虹402。河北:涉县,关克俭5685;武安,关克俭5800;内丘,刘鑫源545;井经,刘继孟3917;平山,1971-07,?497;阜平,刘继孟3262。
  84. 野棉花 图版30: 4-6
  Anemone vitifolia Buch. -Ham. ex DC. Syst. 1: 211. 1817; et Prodr. 1: 21. 1824; Franch. Pl. Delav. 9. 1890; Finet & Gagnep. in Bull. Soc. Bot. France 51: 68. 1904; Ulbr. in Bot. Jahrb. 37: 252. 1905; Pei & Chou, Icon. Chin. Medic. Pl. 1: fig. 15. 1955; W. T. Wang in Fl. Reip. Pop. Sin. 28: 29, pl. 6: 4-6. 1980; in Fl. Xizang. 2: 80. 1985; in Fl. Guangxi 1: 278. 1991; et in Fl. Yunnan. 11: 199, pl 55: 4-6. 2000; Y. K. Li in Fl. Guizhou. 3: 76. 1990; Wang, Ziman & Dutton in Fl. China 6: 317. 2001; Ziman et al. in J. Jap. Bot. 83(3): 141. 2008; L. Xie in Medic. Fl. China 3: 326. 2016. Holotype: Nepal: Lamba, F. Buchuan s. n. (non vidi).
  A. vitifolia var. tomentosa Ulbr. in 1. c. Syntypes: Yunnan, Delavay 2818bis; Sikkim, J. D. Hooker s. n. (non vidi).
  84a. var. vitifolia
  分布于中国云南西部和中部(海拔1 500~2 700 m)、广西西部、贵州南部(1 300 m)、四川(2 400~4 000 m)和西藏(2 030~3 000 m),以及不丹、尼泊尔和印度北部。生于山地草坡,沟边或林中。
  本种的根状茎可供药用,有止咳止血、理气杀虫、祛风除湿之效,用于风湿骨痛、胃寒痛、跌打损伤、蛔虫病、小儿疳积等症;有毒(谢磊等,2016)。
  标本登录 云南:贡山,俞德浚19495;福贡,高黎贡山队27294;大理,苍山,滇西北分队10555;景东,李鸣冈2298;昆明,刘慎谔20230。广西:那坡,中植联广西队4280。贵州:兴仁,张志松,张永田7679。四川:理塘,李文华等81-2041;九寨沟,李文华等82-457。西藏:察隅,王启无65455;通麦,张永田,郎楷永704;昜贡,应俊生,洪德元65-713;墨脱,青藏队1833;隆子,青藏补点队75-542;错那,任再金369;吉隆,姜恕344。
  84b. 绿叶野棉花(变种)
  var. subtomentosa Ulbr. in l. c. Syntypes: Northwestern Himalaya, T. Thomson s. n.; Sikkim, Choongthang, 1903-09, D. Prain s. n. (non vidi).   分布于中國云南(海拔1 350~2 300 m)和西藏东南部(2 000~2 500 m),以及喜马拉雅山区西北部和印度北部。生于山地草坡、林边或林下。
  标本登录 云南:贡山,青藏队9209;德钦,俞德浚10386;福贡,蔡希陶56629;维西,蔡希陶63076;漾濞,刘慎谔22850;楚雄,李鸣冈84;景东,刘炳云53471;澜沧,钱义咏3308;石林,税玉民等64191;屏边,蔡希陶62597;文山,冯国楣11360;砚山,王启无84154;马关,税玉民等31593。
  系3.  块茎银莲花系
  Ser. Anemone. — Sect. Anemone. Tamura in Acta Phytotax. Geobot. 42: 180. 1991; et in Hiepko, Nat. Pflanzenfam., Zwei. Aufl. 17aIV: 347. 1995; Ehrendorfer, Ziman, Konig et al. in Bot. J. Linn. Soc. 160: 312. 2009.
  根状茎块状。基生叶为单叶,3深裂或3全裂,或为三出复叶。聚伞花序简单,具1花,或分枝,具2~3花;总苞苞片(2~)3。萼片5~20。瘦果多数,小,无柄,被绵毛。
  约20种,分布于地中海地区、中亚地区和喜马拉雅山区,以及南美洲和北美洲。中国有1种,分布于新疆西部。
  85. 块茎银莲花 图版12: 4-6
  Anemone gortschakowii  Kar. & Kir, in Bull. Soc. Nat. Mosc. 15: 131. 1842; Juz. in Fl. URSS 7: 260, t. 16: 3. 1937; W. T. Wang in Fl. Reip. Pop. Sin. 28: 34, pl. 8: 4-6. 1980; J. G. Liu in Fl. Xinjiang. 2(1): 275, pl. 74: 1-2. 1994; Wang, Ziman & Dutton in Fl. China 6: 318. 2001. — A. biflora DC, var. gortschakowii (Kar. & Kir.) Sinno in Bot. J. Linn. Soc. 153: 434. 2007; Ehrendorfer, Ziman, Konig et al. in I. c. 160: 327, fig. 5: G. 2009. Described from Alatau, no specimen cited.
  分布于中国新疆西部和西南部以及哈萨克斯坦。生于山地草坡或多石砾山坡,海拔1 400~3 200 m。
  标本登录 新疆:新源,alt. 1 400 m,周太炎65-102;乌恰,alt. 3 200 m,青藏队87-0048。
  一个不了解的杂种:
  Anemone wilsonli Ulbr. hybrid. Nova (A. altaica Fisch. × baicalensis Turcz.) in Bot. Jahrb. 36. Beibl. 80: 2. 1905. Holotype: 湖北西部,1904-05,E. H. Wilson 1853, p. p. (non vidi)。
  从此杂种的形态描述了解到此杂种的花序总苞苞片3枚,有长柄,叶片3~5深裂,这种形态与分布于湖北、四川等省、隶属西南银莲花组(Sect. Anemonanthea)的西南银莲花(A. davidii)的总苞十分相似。此外,还了解到其花序有1~2花,花直径2 cm,萼片5枚,白色,雄蕊花丝丝形,心皮的花柱极短或不存在,柱头无柄或近无柄,上述花序和花的构造与分布于湖北、四川等省、隶属鹅掌草组(Sect. Stolonifera)的鹅掌草(A. flaccida)的花序和花的构造颇为相似。在1905年,E. Ulbrich发现此杂种,至今已过去114年,在此时期中,此变种一直未能第二次被发现,可能已经灭绝。
  新发现的1新种:道孚银莲花,在本文完成后1个月,我在一批标本中又发现银莲花属特产四川西部的1个新种。
  道孚银莲花 (图10)
  Anemonedaofuensis W. T. Wang, sp. nov. Holotype 四川(Sichuan):道孚县,扎拖乡,鲜水河(Daofu Xian, Zhatuo Xiang, Xianshui River), alt. 2 990 m,山坡灌丛,花紫红色(in bushes on slope, fls. purple-red), 2019-04-24,张树仁,李邦泽,林东亮(S. R. Zhang, B. Z. Li & D. L. Lin) 3201 (PE)。
  \[Sect. Himalayicae (Ulbr.) Juz.\]
  Species nova haec est affinis A. obtusilobae D. Don, a qua plantis humilioribus 9-10 cm altis, cymis 1-floris, bracteis involucralibus 2-3-lobatis vel indivisis, sepalis 5, purpureo-rubris, gynoecio ex carpellis 3-4 constanti differt. In A. obtusiloba, plantae 10-30 cm altae, cymae 1(2)-florae, sepala 5 (-8) alba caerulea vel lutea, et gynoecium ex carpellis 8 constans est.   Small perennial herbs, 9-10 cm tall. Basal leaves ca.10, longpetiolate; bladespapery, broad-ovate, 1.1-1.5(-2.4) × 1.6-1.9 (-2.6) cm, at base cordate, 3-sect, central segment broad-rhombic, 3-lobed, with lobes rotundly 1-3-dentate or entire, lateral segments slightly smaller, slightly obliquelly broad-rhombic or subflabellate. 3-lobed, on both surfaces appressed-pubescent; blades of a few small leaves or broad-ovate, 0.5-0.7 × 0.7-1 cm, at base subtruncate, 3-sect, with rhombic2-denticulate central segment and narrow-obovate undivided lateral segments, or broad-rhombic,0.7 × 0.8 cm, at base broadly cuneate, 3-parted; petioles 2.2-4 cm long, pubescent. Scapes 2-3,4-5.5 cm long, densely pubescent, 1-flowered; involucral bracts 3, 0.8-1.1 cm long, pubescent, or one larger cuneate, unequally 3-lobed, two smaller obliquely oblanceolate, equally 2-lobed, or one larger obliquely cuneate, unequally 2-lobed, with the small lobe narrow-ovate, entire and with the larger lobe 3-lobulate; pedicel 4.5-6 cm long, densely appressed-puberulous. Flower: Sepals 5, purple-red (according to field notes), obovate, broad-elliptic or narrow-obovate, 9-12 × 5-7 mm, at apex rounded or obtuse, inconspicuously 7-nerved, adaxially glabrous, abaxially appressed-pubescent. Stamens ca. 30, glabrous; filaments linear, 0.6-1.5 × 0.5 mm; anthers broad-elliptic, 1-1.2 × 1 mm. Carpels 3-4; ovaries ca. 2 mm long, densely pubescent; styles 1-1.2 mm long, glabrous.
  特產于中国四川道孚。生于山坡灌丛中,海拔2 990 m。
  致谢 深切感谢傅德志教授提供银莲花属(Anemone)全部植物名录;谢磊博士、刘冰博士、韩芳桥先生提供文献;班勤、马欣堂二先生提供标本;孙英宝先生为本文绘图,并录入、打印稿件。
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  收稿日期:  2020-07-20
  基金项目:  中国科学院植物研究所资助王文采院士科研活动经费(110100PZ07) [Supported by the Fund for WANG Wen-tsai Academician Scientific Research of Institute of Botany, Chinese Academy of Sciences (110100PZ07)]。
  作者简介: 王文采(1926- ),中国科学院院士,著名植物分类学家,长期从事有花植物分类学研究。
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摘 要: 為探讨国产毛茛科(Ranunculaceae)驴蹄草属(Caltha)两种植物的演化,该文利用传统染色体压片技术和流式细胞术,并结合前人染色体研究结果,对我国驴蹄草23个居群和花葶驴蹄草10个居群进行了细胞学研究。结果表明:驴蹄草是由四倍体(2n=4x=32)、六倍体(2n=6x=48)和八倍体(2n=8x=64)构成的多倍体复合群,花葶驴蹄草具有四倍体(2n=4x=32)和八倍体(2n
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摘要:荷木(Schimasuperba)是鼎湖山南亚热带常绿阔叶林群落中的优势种,为了解南亚热带常绿阔叶林的群落构建机制,该研究分析了鼎湖山南亚热带常绿阔叶林20hm2样地中荷木种群在2005—2017年间的动态变化,并从径级和生境两个方面分析了其死亡率的分布特征。结果表明:(1)荷木种群个体数量从2005—2017年呈现出减少的态势。(2)在径级方面,荷木的死亡率和相对生长率的变化在不同径级区间
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摘 要:田七(Panax notoginseng)是我国的一种传统珍贵的草本药用植物,其重金属污染问题已经引起广泛关注。为了分析田七不同部位对镉毒害的响应,明确不同浓度镉污染对田七体内镉分布的基本特征以及不同部位的富集特性,以揭示镉胁迫对田七不同部位的影响机制及富集转移特性。该研究在“田七之乡”广西靖西市田七园以3年生田七为材料,土培条件下以不施镉处理为空白对照,设置6个镉浓度(5、10、20、3
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摘要:凋落物的可提取腐殖质碳可随着植物生长节律及物候时期的变化而变化,并进而影响物质循环的过程,为深入了解以凋落物为载体的生态系统物质循环特征,该研究以华西雨屏区麻栎(Quercusacutissima)、楠木(Phoebezhennan)、柳杉(Cryptomeriafortune)和喜树(Camptothecaacuminate)为对象,通过定点动态收集萌芽期、展叶期、盛叶期和落叶期的不同类型
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摘 要:杂草给水稻(Oryza sativa)生产带来严重损失,利用水稻自身化感作用被认为是对环境友好的杂草控制方法。稻壳酮A和B是水稻抑制杂草的主要化感物质,其中稻壳酮B是至今发现最高效的天然除草剂之一,具有很好的应用潜力。稻壳酮B可以从水稻根系释放到土壤中抑制周围稗草等植物的种子萌芽和生长。稻壳酮B的浓度大于3 nmol·mL-1时就能抑制水芹和莴苣的根和胚轴生长,同时稻壳酮A和B是水稻重要的
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摘 要:李氏禾(Leersia hexandra)是中國境内发现的第一种铬超积累植物,该文对李氏禾内生菌及其除铬性能进行了研究。采用添加Cr(VI) 的牛肉膏蛋白胨固体平板培养方法,从李氏禾根部分离筛选获得一株具有较强Cr(VI)抗性的内生细菌G04,分子生物学鉴定结果表明该菌株属于阴沟肠杆菌(Enterobacter cloacae)。采用摇瓶培养方法,以Cr(VI)去除率、总Cr(铬)的去除率
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摘要:蚊虫传播疟疾、登革热等多种烈性疾病,严重威胁到人畜健康。特别是生活在西双版纳地区的人们,蚊虫骚扰是长期需要面对的问题。该研究运用民族植物学研究方法,对西双版纳哈尼族驱蚊植物及知识进行调查,共走访了12个哈尼族村寨,访谈91位信息报告人,调查得到24种驱蚊植物,分别属于15科19属。主要利用的部位是植物的叶;燃烧植物烟熏、悬挂放置、楼下铺撒和捣碎涂抹是最为常用的驱蚊方式。通过打分排序法评价出蒿
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摘要:基因组编辑技术的出现对植物遗传育种及作物性状的改良产生了深远意义。CRISPR/Cas(clusteredregularlyinterspacedshortpalindromicrepeat)是由成簇规律间隔短回文重复序列及其关联蛋白组成的免疫系统,其作用是原核生物(40%细菌和90%古细菌)用来抵抗外源遗传物质(噬菌体和病毒)的入侵。该技术实现了对基因组中多个靶基因同时进行编辑,与前两代基
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