Plant cells differ from their animal counterparts such that they are encased in a complex wall structure,crucial for cell growth through a dynamic equilibrium of rigidity and flexibility.Cell wall extensibility can be greatly influenced by apoplastic pH and reactive oxygen species (ROS) (Duan et al.,2014;Haruta et al.,2014).Several recent works demonstrated that the receptor-like kinase (RLK) FERONIA (FER) is critical to fine-tune cell growth in a spatiotemporal and context-dependent manner by controlling apoplastic pH (Haruta et al.,2014) and ROS (Duan et al.,2010,2014),likely balancing wall rigidity for cell integrity and flexibility for cell expansion.