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Diapause is an endocrine-mediated dormancy that occurs at a specific developmental stage (Tauber et al., 1986; Danks, 1987; Kostal, 2006).In the temperate zones, many insects enter a winter diapause and/or summer diapause to escape form the seasonal environmental stresses and survive (Denlinger, 1991; Leather et al., 1993).