The flap-legged and parachute geckos (or flying geckos) of the genera Luperosaurus and Ptychozoon are some of Southeast Asias most rare and enigmatic land vertebrates.The phylogenetic relationships of Luperosaurus,Ptychozoon,and related genera such as Lepidodactylus and Gekko remain unclear.In particular,the phylogenetic placement of the morphologically intermediate taxa Luperosaurus iskandari,and L.gulat are complicated and confusing.Previous results based on mtDNA and one nuclear locus demonstrated that Luperosaurus is more closely related to Lepidodactylus and Pseudogekko than it is to Gekko but that some species currently classified as Luperosaurus are nested within Gekko (Fig.1,Brown et al.,2012).