Patches of ionization are common in the polar ionosphere where their motion and associated density gradients give variable disturbances to High Frequency (HF) radio communications, over-the-horizon radar location errors, and disruption and errors to satellite navigation and communication.Their formation and evolution are poorly understood, particularly under disturbed space weather conditions.We report direct observations of the full evolution of patches during a geomagnetic storm, including formation, polar cap entry, transpolar evolution, polar cap exit, and sunward return flow.Our observations show that modulation of nightside reconnection in the substorm cycle of the magnetosphere helps form the gaps between patches where steady convection would give a "tongue" of ionization (TOI).