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Soybean mosaic virus(SMV)is found in many soybean producing areas,and SMV infection is one of the prevalent viral diseases that can cause significant yield losses in soybean.In plants,resistance(R)genes are involved in pathogen reorganization and innate immune response activation.Most R proteins have nucleotide-binding site and leucine-rich repeat(NBS-LRR)domains,and some of the NBS-LRR type R proteins in dicots have TIR(Toll/Interleukin-1 Receptor)motifs.