论文部分内容阅读
巴戟天花药发育中多糖和脂滴类物质的分布呈现一定的规律:减数分裂之前,花药壁的绒毡层细胞中有少量脂滴,其他细胞中脂滴和淀粉粒都很少。四分体时期,四分体小孢子中开始出现脂滴,绒毡层细胞中的脂滴较以前增加,其他细胞中的脂滴和淀粉粒仍然很少。小孢子早期,游离小孢子在其表面形成了花粉外壁,靠外壁下方有一层周缘分布的多糖物质。绒毡层细胞中的脂滴明显减少。发育晚期的小孢子中形成一个大液泡,细胞质中出现淀粉粒;同时在药壁和药隔组织中也出现了淀粉粒。此时绒毡层退化。在二胞花粉早期,花粉中积累了大量淀粉粒和一些脂滴。但在成熟的花粉中(二胞花粉晚期),淀粉粒消失,只有一定数量的脂滴保留。巴戟天成熟花粉中积累的营养物质主要为脂滴。
The distribution of polysaccharides and lipid droplets during the development of Morinda officinalis showed a certain pattern. Before meiosis, there were a few lipid droplets in the tapetum of anther wall, but few lipid droplets and starch granules in other cells. In the tetrad, lipid droplets began to appear in the tetrad microspores, and the lipid droplets in the tapetum cells increased compared with the previous ones. However, lipid droplets and starch granules in other cells were still scarce. In the early stage of microspore, the free microspores formed the pollen outer wall on the surface of the microspore, and there was a layer of polysaccharide distributed around the outer wall. Lipid droplets in tapetal cells were significantly reduced. The late development of microspores to form a large vacuole, starch granules appear in the cytoplasm; the same time in the drug wall and drug-separated tissue also appeared in the starch granules. The tapetum is degraded at this time. In the early stages of pollen, pollen accumulated large amounts of starch granules and some lipid droplets. However, in mature pollen (late bicellular pollen), starch grains disappear and only a certain amount of lipid droplets remain. Morinda mature pollen accumulation of nutrients mainly lipid droplets.