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掠食性鱼类的捕捞对海洋生态系营养结构的影响现已明确，海洋中鱼类的总产量是由“新生态”硝酸盐的物理可供量决定的，这被称为“上行控制”原理。例如，ｌｖｅｒｓｅｎ（１９９１）指出，上升流中“新生态”硝酸盐的产量与世界海洋鱼类和头足类的年产量之间有很强的相关...
The effect of predatory fish fishing on the nutrient structure of marine ecosystems It is now clear that total fish production in the oceans is determined by the physical availability of “neo-ecological” nitrates, which is known as the “up-control” principle . For example, lversen (1991) pointed out that there is a strong correlation between the yield of “neo-ecologically” nitrate in the ascending stream and the annual output of the world’s marine fish and cephalopods.