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植物木质素是影响纸浆材品质的关键因素,而松科植物作为重要的纸浆材,对其木质素合成相关调控基因进行系统了解,可为完善木质素生物合成模型提供一定的理论依据。笔者通过对国内外已有研究进行分析,总结植物木质素结构、木质素合成途径、木质素合成相关酶基因及松科植物木质素合成基因的研究进展。归纳出松科植物火炬松(Pinus taeda)、马尾松(P.massoniana)、辐射松(P.radiata)、挪威云杉(Picea abie)等木质素合成酶基因研究主要集中在苯丙氨酸解氨酶(phenylalanineammonialyase,PAL)、4-香豆酸辅酶A连接酶(4-coumarate Co A ligase,4CL)、肉桂酸4-羟化酶(cinnamic 4-hydroxygenase,C4H)、香豆酸3-羟化酶(Coumarate 3-hydroxylase,C3H)、咖啡酰辅酶A-O-甲基转移酶(caffeoyl coenzyme A-O-methyltransferase,CCo AOMT)、肉桂酰基辅酶A还原酶(Cinnamoyl Co A reductase,CCR)和肉桂醇脱氢酶(Cinnamyl alcohol dehydrogenase,CAD)等一些常见基因上,其他莽草酸羟基肉桂酰转移酶(shikimate hydroxycinnamoyl transferase,HCT)、阿魏酸5-羟化酶(ferulo 5-hydroxylase,F5H)、O-甲基转移酶(O-methyltransferase,OMT)及咖啡酸-O-甲基转移酶(caffeic acid O-methyltransferase,COMT)等基因研究较欠缺,此外,还有部分基因未涉及。木质素合成是研究松科植物纸浆材的重要切入点,对现有松科植物木质素合成过程中存在问题进行精确的估计,对进一步深入研究松科植物木质素合成机制和优质纸浆材选育具有重要意义。
Plant lignin is the key factor affecting the quality of pulpwood, while Pinaceae, as an important pulp material, has a systematic understanding of its regulatory genes related to lignin synthesis, which may provide some theoretical basis for improving the lignin biosynthesis model. The author analyzes the existing researches both at home and abroad, and summarizes the research progress of plant lignin structure, lignin synthesis pathway, lignin synthesis related enzyme gene and Pinaceae lignin synthesis gene. It is concluded that Pinus taeda, P.massoniana, P.radiata, Picea abie and other lignin synthase genes are mainly concentrated in phenylalanine 4-coumarate Co A ligase (4CL), cinnamic 4-hydroxygenase (C4H), 3-hydroxycoumarinic acid (CMA), cinnamoyl Co A reductase (CCR) and cinnamyl alcohol dehydrogenation (Cinnamyl alcohol dehydrogenase, CAD) and other common genes, other shikimate hydroxycinnamoyl transferase (shikimate hydroxycinnamoyl transferase, ferulo 5-hydroxylase (ferulo 5-hydroxylase, F O-methyltransferase (OMT) and caffeic acid O-methyltransferase (COMT) are not well studied. In addition, some genes are not involved. Lignin synthesis is an important starting point for the study of the pulp material of Pinaceae. The existing problems in the synthesis of Pinus thunbergii Lignin are estimated accurately. For the further study on the synthesis mechanism of lignin and the selection of high-quality pulp material, Significance.