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由于缺少类似于六倍体小麦所拥有的合适细胞遗传原种，因此，限制了四倍体小麦的遗传研究进展。四倍体小麦作为一个种群，很少受到科学家的注意，这是因为四倍体小麦的种植面积只占世界小麦生产总面积的一小部分，而且其面包食品生产中用途有限。然而，有一种四倍体硬粒小麦（ＴｒｉｔｉｃｉｏｎｔｕｒｇｉｄｕｍＬ．ｖａｒｄｕｒｉｏｎ）品种在通心面制品（如ｓｐａｇｈｅｔｔｉ和ｍａｃａｒｏｎｉ）的生产中得到广泛应用。因此，研究重点主要集中在几套硬粒小麦非整倍体遗传原种的生产，以及开发利用这些原种的方法上。硬粒品种Ｌａｎｇｄｏｎ的非整倍体原种包括双－二端体、双单端体、Ｄ－染色体组二体代换系，种间染色体代换系和纯合重组系。
The lack of suitable cytogenetic stock resembles that of hexaploid wheat limits the progress of genetic research on tetraploid wheat. Tetraploid wheat, as a population, seldom receives the attention of scientists because the area planted with tetraploid wheat accounts for only a fraction of the total world wheat production and its use in bakery products is limited. However, there is a species of tetraploid Triticum durum wheat widely used in the production of macaroni products such as spaghetti and macaroni. Therefore, the research focuses mainly on the production of several aneuploid genetic elite strains of durum wheat and the methods of developing and using these elite species. The non-euploid progeny of Langdon, a durum variety, includes two-terminal, two-terminal, two-body D-chromosome substitution lines, interspecific chromosomal substitution lines and homozygous recombinant lines.