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Animals initiate different behaviors in respond to visual stimuli with distinct ecological meanings.Little is known about its underlying neural mechanisms at visuomotor transformation stage though intensive relevant studies have been made at visual processing level.Here, using the visual escape circuit of zebrafish as a model system, we find that dopaminergic neurons in the caudal hypothalamus and specific clusters of glycinergic interneurons in the hindbrain form a functional module, which gates the visuomotor transformation in a stimulus-dependent manner.