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Plants have evolved extraordinarily complex mechanisms to survive pathogen attack.One of these is the hypersensitive response (HR) cell death, a form of programmed cell death (PCD).HRPCD is typically triggered upon recognition of a pathogen-encoded avirulence (Avr) protein by a cognate plant resistance (R) protein.Tobacco N gene is a typical TIR-NB-LRR R gene confers resistance to Tobacco mosaic virus (TMV).