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The maturity locus E1 has a large impact on flowering time in soybean, but the molecular basis for the E1 locus is largely unknown.Through positional cloning, we delimited the E1 locus to a 17.4-kb region containing an intron-free gene (E1).The E1 protein contains a putative bipartite nuclear localization signal and a region distantly related to B3 domain.