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Besides the KU-dependent classical non-homologous end-joining (C-NHEJ) pathway,an alternative NHEJ pathway first identified in mammalian systems, which is often called the back-up NHEJ (B-NHEJ) pathway, was also found in plants.In mammalian systems PARP was found to be one of the essential components in B-NHEJ.Here we investigated whether PARP1 and PARP2 were also involved in B-NHEJ in Arabidopsis.