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長時期來農業上的實際問題之一就是小麥腥黑穗病的研究。這個問題的研究是從各個方面來進行的。初起研究的是黑穗病菌菌種的類別。烏利揚舍夫(1950)的研究闡明了,在蘇聯有三種腥黑穗病菌,即Tilletia tritici wint,T.levis Kuhu,及T.triticoides Savul。最後一種黑穗病菌目前只分佈在阿捷爾拜彊共和國阿爾明尼亞共和國(在外高加索)之間,它與T.tritici wint的區别在於有較小的厚垣抱子。其次研究了春化型的類别。切杰列夫尼柯娃-巴巴揚(1934-1941)的研究確定了在阿爾明尼亞共和國有二個Tilletia tritici wint的春化型:1,f.vulgaris,嚴重感病的麥種爲Triticum vulgaris,中度感病的爲T.persica,T.durum,T.spelta,T.monococcum,T.dicoccum和T.timofeevi,輕度感病的爲T.compactum;2, f.dicoccum,嚴重感病的爲Triticum dicoccum,輕度感病的爲T.vulgaris.在西格魯吉亞還發現了Tilletia tritci wint的第三個春化型,叫做f. timofeevi,只能侵染Triticum timofeevi.
One of the practical problems in agriculture over a long period of time is the study of wheat-based smut. The study of this issue is carried out from all aspects. The earliest studies were the species of the pathogen. Ulyaniusv’s (1950) study clarified that there are three TURs in the Soviet Union, Tilletia tritici wint, T. levis Kuhu, and T.triticoides Savul. The last smut is currently found only in the Azerbaijani Republic of Albania (in the Transcaucasia), which differs from T.tritici wint in that it has a smaller thickness. Second, I studied the categories of vernalization. The study of Cherevnikov-Babayan (1934-1941) identified two vernalizations of Tilletia tritici wint in the Republic of Albania: 1, f.vulgaris, and the severely susceptible wheat species was Triticum vulgaris, moderately susceptible to T. persica, T.durum, T.spelta, T.monococcum, T.dicoccum and T.timofeevi, mildly susceptible to T.compactum; 2, f.dicoccum, The disease is Triticum dicoccum and the mild is T.vulgaris. The third vernalization of Tilletia tritci wint, also known as f. Timofeevi, is found in western Georgia and infects only Triticum timofeevi.