马协不育系系朱英国等用农家品种“马尾粘”的不育株与“协青早选”测交和回交选育成功的。超微结构研究表明马协不育花药单核边位期绒毡层提前解体,液泡膜破裂;不育花粉外壁基粒棒少,且分布不均匀;线粒体有大的电子透明区,二核期不育花粉中央大液泡崩溃。不育花药绒毡层提前解体可能影响花粉外壁合成及花粉营养供应,大量研究表明绒毡层发育异常与花粉败育密切相关,而液泡膜的破坏会导致水解酶类的泄 CMS sterile line Zhu Yingguo and other farmers with “Mawei sticky” sterile lines and “early Qingxuan” test and backcross breeding success. Ultrastructural studies showed that the tapetum at the mononuclear marginal phase of the male sterile anther disintegrated prematurely and the tonoplast ruptured. There were few grana rods on the outer wall of sterile pollen and the distribution was uneven. The mitochondria had a large electron transparent region, Sterile pollen central vacuole collapse. The premature disintegration of sterile anther tapetum may affect the synthesis of pollen exine and the supply of pollen nutrition. A large number of studies have shown that the development of tapetum is closely related to pollen abortion, and the destruction of tonoplast will lead to the release of hydrolase