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H.Morishima(1990)在总结各国近年来对亚洲稻(O.sativa L.)同工酶研究时指出,迄今已发现有20多个同工酶基因座与籼粳稻的分化有关。我们在1990年亦发现一个与亚洲稻起源、演化及分类有关的 Est 基因座,经遗传研究这一基因座上至少有6个复等位基因。分别表现为10A,11A,13A,14A 和15A 条酶带及一条无效酶带,暂时将其命名为 Est_x 基因座,上述6个复等位基因分别命名为 Est_x~(10),Est_x~(11),Est_x~(13),Est_x~(14),Est_x~(15)及 Est_x~0。
H. Morishima (1990) summarized the recent studies on isozymes of O. sativa L. in various countries and pointed out that so far it has been found that more than 20 isozyme loci are involved in the differentiation of indica and japonica. In 1990, we also found an Est locus related to the origin, evolution and classification of rice in Asia. There are at least 6 complex alleles at this genetic locus. Which were respectively expressed as 10A, 11A, 13A, 14A and 15A bands and an inactive enzyme band, tentatively named Est_x locus. The 6 complex alleles were named as Est_x ~ (10), Est_x ~ (11) ), Est_x ~ (13), Est_x ~ (14), Est_x ~ (15) and Est_x ~ 0.