The idea of assembling a Special Issue on Pollen Tube and Reproduction in Molecular Plant was conceived during the organization of the 2012 Pollen Biology Inteational Symposium and Workshop held at the Shanghai Institute of Plant Physiology and Ecology (SIPPE),Chinese Academy of Science (CAS),27-31 October 2012.Wei-Hua Tang of the SIPPE led the organization,with the support of Hong-Wei Xue,Director of SIPPE acting as Chair of the Organization Committee.The 2-d symposium was attended by about 100 inteational participants and focused on reports and discussions on the most recent research activities in the field.The 2-d workshop that followed provided a forum for in-depth introduction on various aspects of pollen and plant reproduction biology to about 200 graduate students,postdoctoral fellows,and new investigators to the field.