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在从事甜菜抗丛根病育种工作中，通过对各类育种材料的抗病性观察，及对其互交和测交后代的调查，发现甜菜的抗丛根病性是由两对显性基因控制的。经过一步分析，提出了甜菜抗丛根病的基因型和遗传模式。假定以“R”和“P”代表这两对显性基因，免疫型为RPRP；抗病型为RPRp、RPrP耐病Ⅰ型为RPrp、RpRp、rPrP；耐病Ⅱ型为Rprp、rPrP；敏感型为rprp。推测：糖用甜菜在驯化过程中，丢失了其中的一对显性基因，假设丢失的“P”，那么，只有耐病Ⅰ型（RpRp）、耐病Ⅱ型（Rprp）和敏感型（rprp）。这样，采用选择的方法，只能选育出耐病型品种，不可能选育出抗病型和免疫型的品种。只有通过与其祖先杂交，将丢失的“P”基因找回来，才能育成抗病型和免疫型品种。
During the breeding of sugarbeet against Rhizobium, we observed the disease resistance of all kinds of breeding materials and the cross-fertilization and cross-fertilization survey. We found that anti-rhizomatosis of sugar beet is composed of two pairs of dominant genes controlling. After a step analysis, put forward beet resistance to rhizomatosis genotypes and genetic patterns. It is assumed that the two pairs of dominant genes are represented by “R” and “P”, the immune type is RPRP, the resistance type is RPRp, the RPrP resistance type I is RPrp, RpRp and rPrP, the resistance type II is Rprp and rPrP, rprp. It is speculated that during the domestication of sugar beet, one pair of dominant genes is lost, and the missing “P” is assumed to have only RpRp, Rprp and rprp. In this way, using the method of selection, we can only breed disease-resistant varieties and it is impossible to breed disease-resistant and immunotype varieties. Only by crossing with their ancestors, the lost “P” gene can be recovered in order to breed disease-resistant and immunotypes.