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曳鳃动物(priapulid)代表了现代动物界一个小门,仅描述了18个现生种,但它们却是寒武纪海洋中最重要的底内生动物,主要化石记录以中国下寒武统帽天山页岩及加拿大中寒武统布尔吉斯页岩为代表.5.3亿年前帽天山页岩中曳鳃动物被划分为6个基本类群:管状蠕虫类(selkirkiid)、棒形虫类(corynetid)、安宁虫类(anningid)、古蠕虫类(palaeoscolecidan)、瘤节虫类(tylotitid)、曳鳃虫类(priapuliid).另外,寒武纪早期另一个重要的底内生蠕形动物类群星虫动物门最近也已报道.中国下寒武统帽天山页岩中曳鳃动物的研究近期取得了一系列研究进展,一些既有属种得到形态修正与深入研究,一些新类型及分类群被描述或建立.其中韩健等研究了帽天山页岩中一类新的古蠕虫化石Tabelliscolex;Huang等修正了帽天山页岩中的Xiaoheiqingella并发表了Yunnanpriapulus,将它们归入曳鳃动物现生科Priapulidaet;Huang等修正了帽天山页岩中曳鳃动物Corynetis,并描述了一个新的类型Anningvermis,分别进行了形态功能学解释,并建立两个绝灭新科Corynetidae和Anningidae;Dong等报道了湖南中晚寒武世直接发育型蠕虫胚胎化石Makueliahunanensis,可能代表了曳鳃动物门古蠕虫类的胚胎发育阶段,韩健等在最近论文中论述了曳鳃形目(Priapulomorpha包括Priapulidae和nibiluchidae)在早寒武世的祖先Xiaoheiqingella和Paratubiluchus.
The priapulid represents a wicket in the modern animal kingdom that describes only 18 present-day species, but they are the most important endophytes in the Cambrian ocean. The main fossil record is the Lower Cambrian Cap Tianshan shale and Canadian Cambrian Shale as the representative. About 530 million years ago, the gill animals in the Tianshan shale were divided into six basic groups: selkirkiid, corynetid, anningid, palaeoscolecidan, Tylotitid, priapuliid, and another important phytoplankton cephalothorax in the early Cambrian has recently been reported. Recently, a series of research progresses have been made in the study of the triassic goby in the Lower Cambrian capped Tianshan shale in China. Some existing species have been subjected to morphological modification and further study, and some new types and taxa have been described or established. Among them, Han et al. Studied a new type of ancient worm fossil, Tabelliscolex, in the capped Tianshan shale; Huang et al amended the Xiaoheiqingella in the capped Tianshan shale and published Yunnanpriapulus and classified them into Priapulidaet, Revised Corynetis in the Tianshan shale, and described a new type of Anningvermis, which were morphologically functionalized and two new extinctions Corynetidae and Anningidae were established respectively. Dong et al. Reported that the Late-Cambrian The direct developmental worm embryo fossil Makuelia hununii may represent the stage of embryonic development of the ancient worms in the phylum Trachinus. In recent papers, Han Jian et al. Discussed the ancestors of Priapulomorpha including Priapulidae and nibiluchidae in the Early Cambrian Xiaoheiqingella and Paratubiluchus.