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对取自法国不同地区的葡萄霜霉菌群体的孢子囊的观察表明,该菌致病力的差异首先表现在其孢子囊的大小和其中所含有的细胞核数量的差异。通过连续孢子囊移种,由任意取自自然群体的一个孢子囊获得了几个致病力各异的纯系。致病力不同的纯系混合侵染,可提高混合群体的致病力,利用对 Metalaxyl的抗性作标记,证明了不同菌丝体间的细胞核交换现象和异核现象。由于异核现象的作用,霜霉菌的自然群体的致病力可以通过细胞核之间的协生作用达到其最高水平。
Observations on the sporocysts of grape downy mildew populations taken from different regions of France showed that the difference in virulence of the bacteria was first shown by the difference in the size of the sporocysts and the number of nuclei contained therein. By continuous sporangiobservation, several virulent lines were obtained from one sporangia arbitrarily taken from a natural population. The virulent mixed infection of different pathogenicity can increase the virulence of the mixed population and mark the resistance of Metalaxyl to prove the nuclear exchange phenomenon and heteronuclear phenomenon among different mycelium. Due to heterotrophic effects, the virulence of the natural populations of downy mildew can reach their highest levels through synergy between the nuclei.