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Cell locomotion is essential for early development,angiogenesis,tissue regeneration,the immune response,andwound healing in multicellular organisms,and plays a verydeleterious role in cancer metastasis in humans.Locomotioninvolves the detection andtransduction of extracellular chemical and mechanical signals,integration of thesignals into an intracellular signal,and the spatio-temporal control of the intracellularbiochemical and mechanical responses that lead to force generation,morphologicalchanges and directed movement.