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Tomato fruit-weight 2.2 (FW2.2) was reported to control up to 30% fruit weight.Recent studies demonstrated that FW2.2-1ike (FWL) genes also play important roles in plant growth and development.For instance, a maize homolog of FW2.2, named cell number regulator 1 (CNR1), negatively regulates plant and organ size.However, FWL genes in rice have not been characterized yet.