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Cytosine DNA methylation is a stable epigenetic mark that is frequently associated with the silencing of genes and transposable elements (TEs).In Arabidopsis, the establishment of DNA methylation is through the RNA-directed DNA methylation (RdDM) pathway.To identify components of the RdDM machinery, we carried out a forward genetic screen on a T-DNA-mutagenized population in the ros1 background, based on reactivation of the RD29A promoter-driven luciferase reporter gene.