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The CF-FM signal emitted by CF-FM bats may be called "a simple-complex sound signal",which CF and FM components have different function during echolocation(O‘Neill & Suga,1982; Kober et al.,1990; Fu et al.,2010; Schnitzler & Denzinger,2011; Jakobsen et al.,2013).How to process the complex and behavior-related sound signals in the auditory system is known very little up to now.