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Endoplasmic reticulum(ER)plays important roles in viral replication.Viral infection can lead to endoplasmic reticulum stress response(ER stress),The ER stress is marked by the activation of a series of signaling pathways,called the unfolded protein response(UPR),which consists of three distinct signal pathways,PRKR-like ER kinase(PERK),inositol-requiring enzyme 1(IRE 1)and activated transcription factor 6(ATF6).