晚石炭世莫斯科期刺毛类(Chaetetids)是重要的造礁生物,在全球范围内广泛分布并建造礁体。中国已报道的刺毛类礁主要分布在辽东太子河流域,产在上石炭统本溪组海相灰岩中。刺毛类生长形态多样,有层状、板状、丘状、分枝状和柱状,主要以板状覆盖沉积物的方式造礁。研究区刺毛类礁中刺毛类生长形态及其所主导的生物群落在垂向上变化明显,由此将礁体的建造划分为三个阶段:(1)定殖阶段:刺毛类较少,生长方式单一,多以层状形态生长在棘屑滩上。层状刺毛类体积小,抗浪能力较弱,很难在其周围创造水流平缓的可供其他底栖生物生存的环境。因而该阶段群落生物种类单一,数量稀少;(2)繁盛阶段:刺毛类丰富,生长形态多样,以板状、分枝状和丘状为主。板状刺毛类单独形成的礁骨架,或与分枝状和丘状刺毛类组合起来的礁骨架,都能在其周围一定范围营造适宜底栖生物生存的环境,因此该阶段群落的生物多样性高;(3)衰退阶段:刺毛类很少,多为薄层状,断续生长在棘屑组成的基底上。薄层状刺毛类很脆弱,基本失去改善周围环境的能力,因此该阶段群落生物多样性很低。以上分析说明刺毛类生长形态的变化可以影响生物群落的组成和演化。生物群落的演化过程就是礁体建造的过程。从刺毛类生长形态的角度研究礁体建造过程,为分析由具有多种生长形态的造礁生物建造的生物礁的成礁机制提供新思路。 Late Carboniferous The Moscow Period The Chaetetids are important reef creatures that are widely distributed and built around the world. China has reported that the barren reefs are mainly distributed in the Taizi River basin in Liaodong and are produced in the marine limestone of the Upper Carboniferous Benxi Formation. Bristles species growth and diversity, layered, plate-shaped, mound-shaped, branched and columnar, mainly in the form of plate-covered sediment reef. In the study area, the growth pattern of the britain and the dominant biota in the brittle reef varied greatly in the vertical direction. Therefore, the construction of the reef was divided into three stages: (1) colonization stage: less brittle bristles , A single mode of growth, and more layered growth in the echinoderms beach. Layered bristles are small in size and weak in wave resistance, making it difficult to create an environment around them that allows for smooth flow of other benthic organisms. Therefore, the species of community was single and the number was scarce at this stage. (2) Prosperous stage: The species of bristles were rich and the growth patterns were diverse. The main types were plate, branch and mound. Reef skeletons formed by plate-shaped brier types alone or reef skeletons combined with branch-shaped and moulted tuberous species can create a suitable environment for benthic organisms to survive in a certain range around them. Therefore, High diversity; (3) decline stage: brittle little, mostly lamellar, intermittent growth in the composition of the substrate on the chip. Thin layer of brittle brittle, basically lost the ability to improve the surrounding environment, so the stage of biological diversity is very low. The above analysis shows that changes in the growth pattern of the brittle species can affect the composition and evolution of the biome. The evolution of biomes is the process of reef construction. The reconstruction process of reefs was studied from the perspective of the growth patterns of brittle barks and a new idea was provided to analyze the reef mechanism of reefs constructed by reef-building organisms with various growth patterns.